A midge is any small

fly Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwing ...

, including species in several

families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...

of non-

mosquito Mosquitoes (or mosquitos) are members of a group of almost 3,600 species of small flies within the family Culicidae (from the Latin ''culex'' meaning " gnat"). The word "mosquito" (formed by ''mosca'' and diminutive ''-ito'') is Spanish for "li ...

Nematocera The Nematocera (the name means "thread-horns") are a suborder of elongated flies with thin, segmented antennae and mostly aquatic larvae. This group is paraphyletic and contains all flies but species from suborder Brachycera (the name means "sh ...

Diptera Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...

. Midges are found (seasonally or otherwise) on practically every land area outside permanently arid deserts and the frigid zones. Some midges, such as many

Phlebotominae The Phlebotominae are a subfamily of the family Psychodidae. In several countries, their common name is sandfly; but that name is also applied to other flies. The Phlebotominae include many genera of blood-feeding (hematophagous) flies, including ...

(sand fly) and Simuliidae (black fly), are vectors of various diseases. Many others play useful roles as prey for

insectivore A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores wer ...

s, such as various

frogs A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-frog" ''Triadobatrachus'' is ...

and

swallows The swallows, martins, and saw-wings, or Hirundinidae, are a family of passerine songbirds found around the world on all continents, including occasionally in Antarctica. Highly adapted to aerial feeding, they have a distinctive appearance. The ...

. Others are important as

detritivore Detritivores (also known as detrivores, detritophages, detritus feeders, or detritus eaters) are heterotrophs that obtain nutrients by consuming detritus (decomposing plant and animal parts as well as feces). There are many kinds of invertebrates, ...

s, and form part of various

nutrient cycles A nutrient cycle (or ecological recycling) is the movement and exchange of inorganic and organic matter back into the production of matter. Energy flow is a unidirectional and noncyclic pathway, whereas the movement of mineral nutrients is cycli ...

. The habits of midges vary greatly from species to species, though within any particular family, midges commonly have similar ecological roles. Examples of families that include species of midges include: *

Blephariceridae The Blephariceridae, commonly known as net-winged midges, are a nematoceran family (biology), family in the order Fly, Diptera. The adults resemble tipulidae, crane flies except with a projecting anal angle in the wings, and different head shape, ...

, net-winged midges *

Cecidomyiidae Cecidomyiidae is a family of flies known as gall midges or gall gnats. As the name implies, the larvae of most gall midges feed within plant tissue, creating abnormal plant growths called galls. Cecidomyiidae are very fragile small insects usu ...

, gall midges *

Ceratopogonidae Ceratopogonidae is a family of flies commonly known as no-see-ums, or biting midges, generally in length. The family includes more than 5,000 species, distributed worldwide, apart from the Antarctic and the Arctic. Ceratopogonidae are holomet ...

, biting midges (also known as no-see-ums or punkies in

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

br>BugGuide
/ref> and sandflies in Australia) *

Chaoboridae Chaoboridae, commonly known as phantom midges or glassworms, is a family of fairly common midges with a cosmopolitan distribution. They are closely related to the Corethrellidae and Chironomidae; the adults are differentiated through peculiarit ...

, phantom midges *

Chironomidae The Chironomidae (informally known as chironomids, nonbiting midges, or lake flies) comprise a family of nematoceran flies with a global distribution. They are closely related to the Ceratopogonidae, Simuliidae, and Thaumaleidae. Many species ...

, non-biting midges (also known as muckleheads, muffleheads or lake flies in the

Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes in the mid-east region of North America that connect to the Atlantic Ocean via the Saint Lawrence River. There are five lakes ...

region of

) *

Deuterophlebiidae The fly genus ''Deuterophlebia'' is the sole member of the small monogeneric family Deuterophlebiidae or mountain midges. Adults have broad, fan-shaped wings, and males have extremely long antennae which they employ when contesting territories ov ...

, mountain midges *

Dixidae The Dixidae (meniscus midges) are a family (biology), family of aquatic insect, aquatic nematoceran fly, flies (Diptera). The larvae live in unpolluted, standing fresh waters, just beneath the surface film, usually amongst marginal aquatic vegetat ...

, meniscus midges *

Scatopsidae __NOTOC__ The minute black scavenger flies or "dung midges", are a family, Scatopsidae, of nematoceran flies. Despite being distributed throughout the world, they form a small family with only around 250 described species in 27 genera, although m ...

, dung midges *

Thaumaleidae Thaumaleidae, the solitary midges or trickle midges, are a group of nematoceran flies related to the Ceratopogonidae, Chironomidae, and the Simuliidae. They are small, stocky, yellow to brown flies (3–4 mm). Very few species are known fo ...

, solitary midges

Examples

The

(biting midges) include serious blood-sucking pests, feeding both on humans and other mammals. Some of them spread the livestock diseases blue tongue and

African horse sickness African horse sickness (AHS) is a highly infectious and deadly disease caused by ''African horse sickness virus''. It commonly affects horses, mules, and donkeys. It is caused by a virus of the genus ''Orbivirus'' belonging to the family ''Reovir ...

– other species though, are at least partly nectar feeders, and some even suck insect bodily fluids. Ceratopogonidae midge sucking Sphodromantis blood IMG 3045ss

Ceratopogonidae midge sucking Sphodromantis blood IMG 3045ss

Other ceratopogonid midges are major pollinators of ''

Theobroma cacao ''Theobroma cacao'', also called the cacao tree and the cocoa tree, is a small ( tall) evergreen tree in the family Malvaceae. Its seeds, cocoa beans, are used to make chocolate liquor, cocoa solids, cocoa butter and chocolate. The largest pro ...

'' (cocoa tree). Having natural pollinators has beneficial effects in both agricultural and biological products because it increases crop yield and also density of predators of the midges (still beneficial to all parties). Most other midge families are bloodsuckers, but it is not possible to generalise rigidly because of the vagueness of the term "midge". There is, for example, no objective basis for excluding the

Psychodidae Psychodidae, called drain flies, sink flies, filter flies, sewer flies, or sewer gnats, is a family of true flies. Some genera have short, hairy bodies and wings giving them a "furry" moth-like appearance, hence one of their common names, moth f ...

from the list, and some of them (or midge-like taxa commonly included in the family, such as ''

Phlebotomus ''Phlebotomus'' is a genus of " sand flies" in the Diptera family Psychodidae. In the past, they have sometimes been considered to belong in a separate family, Phlebotomidae, but this alternative classification has not gained wide acceptance. E ...

'') are blood-sucking pests and disease vectors. Most midges, apart from the gall midges (

), are aquatic during the larval stage. Some Cecidomyiidae (e.g., the sorghum midge) are significant plant pests. The larvae of some

contain

hemoglobin Hemoglobin (haemoglobin BrE) (from the Greek word αἷμα, ''haîma'' 'blood' + Latin ''globus'' 'ball, sphere' + ''-in'') (), abbreviated Hb or Hgb, is the iron-containing oxygen-transport metalloprotein present in red blood cells (erythrocyte ...

and are sometimes referred to as bloodworms. Non-biting midge flies are a common minor nuisance around artificially-created bodies of water.

Examples

See also

References

Further reading