''Grusonia pulchella'' (Engelm.) H.Rob., also known as sagebrush cholla, is a tuberous

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of opuntioid

cactus A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Greek ...

from the

Mojave Desert The Mojave Desert ( ; mov, Hayikwiir Mat'aar; es, Desierto de Mojave) is a desert in the rain shadow of the Sierra Nevada mountains in the Southwestern United States. It is named for the indigenous Mojave people. It is located primarily in ...

of central

Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...

, eastern

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

, northwestern

Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...

and western

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...

in the United States. ''Grusonia pulchella'' has at various times been included in ''

Opuntia ''Opuntia'', commonly called prickly pear or pear cactus, is a genus of flowering plants in the cactus family Cactaceae. Prickly pears are also known as ''tuna'' (fruit), ''sabra'', ''nopal'' (paddle, plural ''nopales'') from the Nahuatl word f ...

'' or placed in a separate genus ''Micropuntia''.

Habitat

''Grusonia pulchella'' grows in gravelly

alluvial fan An alluvial fan is an accumulation of sediments that fans outwards from a concentrated source of sediments, such as a narrow canyon emerging from an escarpment. They are characteristic of mountainous terrain in arid to semiarid climates, but a ...

s, often above salt flats or

alkali In chemistry, an alkali (; from ar, القلوي, al-qaly, lit=ashes of the saltwort) is a basic, ionic salt of an alkali metal or an alkaline earth metal. An alkali can also be defined as a base that dissolves in water. A solution of a ...

basins. Specimens can be locally common, although they are difficult to locate, often growing under other shrubs such as

shadscale ''Atriplex confertifolia'', the shadscale or spiny saltbush, is a species of evergreen shrub in the family Amaranthaceae, which is native to the western United States and northern Mexico. Description The height of ''Atriplex confertifolia'' vari ...

Description

''Grusonia pulchella'' differs from other North American opuntioid cacti in having a

geophytic A storage organ is a part of a plant specifically modified for storage of energy (generally in the form of carbohydrates) or water. Storage organs often grow underground, where they are better protected from attack by herbivores. Plants that have ...

habit, where above-ground growth dies back to the crown in adverse conditions, and resprouts under more favorable conditions. Authors have described the underground storage structure as a "tuberous root" or true

tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...

. The above-ground stems are variable, being cylindrical to globular. The areoles bear flexible yellow spines and white wool. Overall, the above-ground growth can often form a cushion-plant habit. The flowers are bright magenta, and the fruits are dehiscent. The seeds are unique, with a groove running along the hilar surface.Flora of North America, vol 4. 2004.

References

Bibliography

* {{Taxonbar, from1=Q1551117, from2=Q14954472 pulchella Cacti of the United States Flora of the California desert regions Flora of the Great Basin Flora of Arizona Flora of California Flora of Nevada Flora of Utah Natural history of the Mojave Desert Flora without expected TNC conservation status