Microecosystems can exist in locations which are precisely defined by critical
environmental factors within small or tiny spaces.
Such factors may include
temperature
Temperature is a physical quantity that quantitatively expresses the attribute of hotness or coldness. Temperature is measurement, measured with a thermometer. It reflects the average kinetic energy of the vibrating and colliding atoms making ...
,
pH,
chemical
A chemical substance is a unique form of matter with constant chemical composition and characteristic properties. Chemical substances may take the form of a single element or chemical compounds. If two or more chemical substances can be combin ...
milieu,
nutrient
A nutrient is a substance used by an organism to survive, grow and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...
supply, presence of symbionts or solid substrates, gaseous atmosphere (
aerobic or
anaerobic
Anaerobic means "living, active, occurring, or existing in the absence of free oxygen", as opposed to aerobic which means "living, active, or occurring only in the presence of oxygen." Anaerobic may also refer to:
*Adhesive#Anaerobic, Anaerobic ad ...
) etc.
Some examples
Pond microecosystems
These microecosystems with limited
water
Water is an inorganic compound with the chemical formula . It is a transparent, tasteless, odorless, and Color of water, nearly colorless chemical substance. It is the main constituent of Earth's hydrosphere and the fluids of all known liv ...
volume are often only of temporary duration and hence colonized by organisms which possess a drought-resistant
spore
In biology, a spore is a unit of sexual reproduction, sexual (in fungi) or asexual reproduction that may be adapted for biological dispersal, dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores fo ...
stage in the lifecycle, or by organisms which do not need to live in water continuously. The ecosystem conditions applying at a typical pond edge can be quite different from those further from shore. Extremely space-limited water ecosystems can be found in, for example, the water collected in
bromeliad
The Bromeliaceae (the bromeliads) are a Family (biology), family of monocot flowering plants of about 80 genera and 3700 known species, native mainly to the Tropics, tropical Americas, with several species found in the American subtropics and on ...
leaf bases and the "pitchers" of ''
Nepenthes
''Nepenthes'' ( ) is a genus of carnivorous plants, also known as tropical pitcher plants, or monkey cups, in the monotypic family Nepenthaceae. The genus includes about 170 species, and numerous natural and many cultivated hybrids. They are m ...
''.
Animal gut microecosystems
These include the
buccal region (especially cavities in the
gingiva
The gums or gingiva (: gingivae) consist of the mucosal tissue that lies over the mandible and maxilla inside the mouth. Gum health and disease can have an effect on general health.
Structure
The gums are part of the soft tissue lining of the ...
),
rumen
The rumen, also known as a paunch, is the largest stomach compartment in ruminants. The rumen and the reticulum make up the reticulorumen in ruminant animals. The diverse microbial communities in the rumen allows it to serve as the primary si ...
,
caecum
The cecum ( caecum, ; plural ceca or caeca, ) is a pouch within the peritoneum that is considered to be the beginning of the large intestine. It is typically located on the right side of the body (the same side of the body as the appendix, ...
etc. of
mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
ian
herbivores
A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...
or even
invertebrate
Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
digestive tract
The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the Digestion, digestive system that leads from the mouth to the anus. The tract is the largest of the body's systems, after the cardiovascula ...
s. In the case of mammalian gastrointestinal microecology, microorganisms such as protozoa, bacteria, as well as curious incompletely defined organisms (such as certain large structurally complex
Selenomonads, ''Quinella ovalis'' "Quin's Oval", ''Magnoovum eadii'' "Eadie's Oval", ''Oscillospira'' etc.) can exist in the rumen as incredibly complex, highly enriched mixed populations, (see Moir and Masson images
). This type of microecosystem can adjust rapidly to changes in the nutrition or health of the host animal (usually a ruminant such as cow, sheep, goat etc.); see Hungate's "The Rumen and its microbes 1966). Even within a small closed system such as the rumen there may exist a range of ecological conditions: Many organisms live freely in the rumen fluid whereas others require the substrate and metabolic products supplied by the stomach wall tissue with its folds and interstices. Interesting questions are also posed concerning the transfer of the strict
anaerobe
An anaerobic organism or anaerobe is any organism that does not require molecular oxygen for growth. It may react negatively or even die if free oxygen is present. In contrast, an aerobic organism (aerobe) is an organism that requires an oxygenat ...
organisms in the gut microflora/microfauna to the next host generation. Here, mutual licking and
coprophagia
Coprophagia ( ) or coprophagy ( ) is the consumption of feces. The word is derived from the Ancient Greek "feces" and "to eat". Coprophagy refers to many kinds of feces-eating, including eating feces of other species (heterospecifics), of o ...
certainly play important roles.
Soil microecosystems
A typical soil microecosystem may be restricted to less than a millimeter in its total depth range owing to steep variation in humidity and/or atmospheric gas composition. The soil grain size and physical and chemical properties of the substrate may also play important roles. Because of the predominant solid phase in these systems they are notoriously difficult to study microscopically without simultaneously disrupting the fine spatial distribution of their components.
Terrestrial hot-spring microecosystems
These are defined by gradients of water temperature, nutrients, dissolved gases,
salt
In common usage, salt is a mineral composed primarily of sodium chloride (NaCl). When used in food, especially in granulated form, it is more formally called table salt. In the form of a natural crystalline mineral, salt is also known as r ...
concentrations etc. Along the path of terrestrial water flow the resulting temperature gradient continuum alone may provide many different minute microecosystems, starting with thermophilic bacteria such as
Archaea
Archaea ( ) is a Domain (biology), domain of organisms. Traditionally, Archaea only included its Prokaryote, prokaryotic members, but this has since been found to be paraphyletic, as eukaryotes are known to have evolved from archaea. Even thou ...
"Archaebacteria" ( or more), followed by conventional
thermophiles (),
cyanobacteria
Cyanobacteria ( ) are a group of autotrophic gram-negative bacteria that can obtain biological energy via oxygenic photosynthesis. The name "cyanobacteria" () refers to their bluish green (cyan) color, which forms the basis of cyanobacteri ...
(blue-green algae) such as the motile filaments of ''Oscillatoria'' (),
protozoa
Protozoa (: protozoan or protozoon; alternative plural: protozoans) are a polyphyletic group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic debris. Historically ...
such as Amoeba,
rotifers
The rotifers (, from Latin 'wheel' and 'bearing'), sometimes called wheel animals or wheel animalcules, make up a phylum (Rotifera ) of microscopic and near-microscopic pseudocoelomate animals.
They were first described by Rev. John Harris ...
, then
green algae
The green algae (: green alga) are a group of chlorophyll-containing autotrophic eukaryotes consisting of the phylum Prasinodermophyta and its unnamed sister group that contains the Chlorophyta and Charophyta/ Streptophyta. The land plants ...
() etc.
Of course other factors than temperature also play important roles. Hot springs can provide classic and straightforward ecosystems for microecology studies as well as providing a haven for hitherto undescribed organisms.
Deep-sea microecosystems
The best known contain rare specialized organisms, found only in the immediate vicinity (sometimes within centimeters) of underwater
volcanic vent
A volcano is commonly defined as a vent or fissure in the crust of a planetary-mass object, such as Earth, that allows hot lava, volcanic ash, and gases to escape from a magma chamber below the surface.
On Earth, volcanoes are most often fo ...
s (or "smokers"). These ecosystems require extremely advanced diving and collection techniques for their scientific exploration.
Closed microecosystem
One that is sealed and completely independent of outside factors, except for temperature and light. A good example would be a plant contained in a sealed jar and submerged under water. No new factors would be able to enter this ecosystem.
References
{{modelling ecosystems, expanded=other
Ecosystems
Environmental science
Ecology