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Metriorhynchoidea is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
of
thalattosuchia Thalattosuchia is a clade of marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of Cro ...
n
crocodyliform Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...
s from the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
to the
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
(
Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarcian ...
-
Valanginian In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 139.8 ± 3.0 Ma and 132.9 ± 2.0 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretaceou ...
, possibly as late as early
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
) of
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
. Metriorhynchids are fully aquatic
crocodyliform Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...
s. Named by Fitzinger, in 1843, it contains the basal taxa like ''
Teleidosaurus ''Teleidosaurus'' is an extinct genus of carnivorous metriorhynchoid crocodyliform from Middle Jurassic (late Bajocian to early Bathonian stage) deposits of Normandy, France.Wilkinson LE, Young MT, Benton MJ. 2008. A new metriorhynchid crocodilia ...
'', ''
Zoneait ''Zoneait'' (pronounced "zone-eight" and meaning "large tooth" in the Kiowa language) is an extinct genus of thalattosuchian crocodylomorph known from a single species, ''Zoneait nargorum'', from the Middle Jurassic of Oregon. ''Z. nargorum'' was ...
'' and ''
Eoneustes ''Eoneustes'' (meaning "dawn swimmer") is an extinct genus of metriorhynchoid crocodyliform from Middle Jurassic (late Bajocian to early Bathonian stage) deposits of France. ''Eoneustes'' was a carnivore that lived in the oceans and spent much, ...
'' and the family
Metriorhynchidae Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period (Bajocian to early Aptian) of Europe, North America and South America. The name Metriorhync ...
. An unnamed taxon is known from
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
.


Phylogeny

Metriorhynchoidea is a
stem-based taxon Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional approach, in which taxon names are defined by a '' type'', which ...
defined in 2009 as the most inclusive clade consisting of ''
Metriorhynchus geoffroyii ''Metriorhynchus'' is an extinct genus of marine crocodyliform that lived in the oceans during the Late Jurassic. The type species, ''M. brevirostris'' was named in 1829 as a species of ''Steneosaurus'' before being named as a separate genus by ...
'', but not ''
Teleosaurus cadomensis ''Teleosaurus'' (from el, τέλειος , 'perfect' and el, σαῦρος , 'lizard') is an extinct genus of teleosaurid crocodyliform found in the Middle Jurassic Calcaire de Caen Formation of France. It was approximately in length and weig ...
''. The
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
below follows the topology from a 2011 analysis by Andrea Cau and Federico Fanti. Note that the same topology was obtained in Mark T. Young and Marco Brandalise de Andrade, 2009 and Mark T. Young, Stephen L. Brusatte, Marcello Ruta and Marco Brandalise de Andrade, 2010.


References

Middle Jurassic crocodylomorphs Early Cretaceous crocodylomorphs Vertebrate superfamilies Prehistoric marine crocodylomorphs Late Jurassic crocodylomorphs Early Jurassic first appearances Early Cretaceous extinctions Thalattosuchians {{paleo-archosaur-stub