Messelasturidae
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Messelasturidae is an extinct family of birds known from the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
of North America and Europe. Their morphology is a mosiac that in some aspects are strongly convergent with modern
hawks Hawks are birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. This subfamily a ...
and
falcons Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Falcons are widely distributed on all continents of the world except Antarctica, though closely related raptors did occur there in the Eocene. Adult falcons ...
, but in others are more similar to
parrot Parrots, also known as psittacines (), are birds of the roughly 398 species in 92 genera comprising the order Psittaciformes (), found mostly in tropical and subtropical regions. The order is subdivided into three superfamilies: the Psittacoid ...
s. Initially interpreted as stem-
owls Owls are birds from the Order (biology), order Strigiformes (), which includes over 200 species of mostly Solitary animal, solitary and Nocturnal animal, nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vi ...
, more recent studies have suggested a closer relationship to parrots and
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
s. Their ecology is enigmatic. The known messelasturid species are ''Messelastur gratulator'', ''Tynskya eocaena'', and ''T. waltonensis''.


Description

On the skull, Messelasturidae possessed a
raptorial The term ''raptorial'' implies much the same as ''predatory'' but most often refers to modifications of an arthropod's foreleg that make it function for the grasping of prey while it is consumed, where the gripping surfaces are formed from the o ...
beak and a large supraorbital process. On the feet, messelasturids lacked an ossified supratendineal bridge on the distal
tibiotarsus The tibiotarsus is the large bone between the femur and the tarsometatarsus in the leg of a bird. It is the fusion of the proximal part of the tarsus with the tibia. A similar structure also occurred in the Mesozoic Heterodontosauridae. These sm ...
. They also featured raptorlike
ungual An ungual (from Latin ''unguis'', i.e. ''nail'') is a highly modified distal toe bone which ends in a hoof, claw, or nail. Elephants and ungulates have ungual phalanges, as did the sauropods and horned dinosaurs. A claw is a highly modified ungual ...
phalanges The phalanges (singular: ''phalanx'' ) are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bones. ...
. They were likely ground dwelling carnivores. They existed at an age range of 61.7 to 40.4 Ma. ''Tynskya'' is characterized by a distinctive morphology of the
tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) and meta ...
.


Discovery and classification

'' Messelastur gratulator'' was originally described in 1994 from two skulls with associated vertebrae, found in the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
aged
Messel pit The Messel pit (german: Grube Messel) is a disused quarry near the village of Messel ( Landkreis Darmstadt-Dieburg, Hesse) about southeast of Frankfurt am Main, Germany. Bituminous shale was mined there. Because of its abundance of well-preserv ...
, Germany. ''
Tynskya eocaena ''Tynskya'' is a genus of messelasturid bird. It is known from a fossil of the North American Green River Formation and the London Clay Formation of England, both from the early Eocene The Eocene ( ) Epoch is a geological epoch that lasted ...
'' was described from specimens found in the Eocene aged
Green River Formation The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes in three basins along the present-day Green River in Colorado, Wyoming, and Utah. The sediments are deposited in very fine ...
in the western United States by
Gerald Mayr Gerald Mayr is a German palaeontologist who is Curator of Ornithology at the Senckenberg Research Institute in Frankfurt am Main, Hesse. He has published extensively on fossil birds, especially the Paleogene avifauna of Europe. He is an expert on ...
in 2000, with indeterminate similar remains also being noted from the equivalently aged
London Clay The London Clay Formation is a marine geological formation of Ypresian (early Eocene Epoch, c. 56–49 million years ago) age which crops out in the southeast of England. The London Clay is well known for its fossil content. The fossils from t ...
of southern England. In 2004, a more complete skeleton of ''Messelastur'' was described, and it and ''Tynskya'' were placed into their own family Messelasturidae by Mayr, as the sister group of Strigiformes or possibly
Falconiformes The order Falconiformes () is represented by the extant family Falconidae (falcons and caracaras) and a handful of enigmatic Paleogene species. Traditionally, the other bird of prey families Cathartidae (New World vultures and condors), Sagitt ...
. An additional specimen of ''Messelastur'' from Messel pit was described in 2011 by Mayr, finding that Messelasturidae was closely related to the extinct family
Halcyornithidae Halcyornithidae or Pseudasturidae is a family of fossil birds, possibly belonging to the order Psittaciformes (parrots and relatives). Members of this family lived in the Eocene, around 55-48 million years ago. Fossil remains are known from the ...
, and both were likely
stem-group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. ...
s to Psittaciformes. A study in 2021 described a new species of ''Tynskya'', ''T. waltonensis'' from the London Clay. A phylogenetic analysis constrained to molecular data conducted in the study found messelasturids closely related to
Psittacopasserae Psittacopasserae is a taxon of birds consisting of the Passeriformes (passerines, a large group of perching birds) and Psittaciformes (parrots). Per Ericson and colleagues, in analysing genomic DNA, revealed a lineage comprising passerines, psi ...
, the clade containing
Passeriformes A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
and Psittaciformes, but a sister group relationship with Halcyornithidae was not recovered.


Ecology

Due to the mosaic nature of the morphology of messelasturids, their ecology is difficult to assess. It is likely they had a specialised feeding ecology. It is unlikely that they regularly searched the ground for food or engaged in
hawking Hawking may refer to: People * Stephen Hawking (1942–2018), English theoretical physicist and cosmologist * Hawking (surname), a family name (including a list of other persons with the name) Film * ''Hawking'' (2004 film), about Stephen Ha ...
via sallying flights from perches. It is possible that they had a raptorial ecology, and hard shelled invertebrates have also been suggested as a food source, but other items like fruit and seeds cannot be ruled out.


Distribution

Fossils of ''Messelastur'' are known from Germany (Messel). ''Tynskya'' remains have been found in the US (
Green River Formation The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes in three basins along the present-day Green River in Colorado, Wyoming, and Utah. The sediments are deposited in very fine ...
), and in the UK (
London Clay The London Clay Formation is a marine geological formation of Ypresian (early Eocene Epoch, c. 56–49 million years ago) age which crops out in the southeast of England. The London Clay is well known for its fossil content. The fossils from t ...
), illustrating the resemblance between the early Eocene birds of North America and Europe. Indeterminate messelasturid remains have also been reported from the early Eocene
Nanjemoy Formation The Nanjemoy Formation is a geologic formation pertaining to both the Wilcox Group and the Pamunkey Group of the eastern United States, stretching across the states of Virginia, Maryland, and District of Columbia. The formation crops out east of ...
of Virginia in the United States. An indeterminate bird from the
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
aged
Menat In ancient Egyptian religion, a menat ( egy, mnj.t, ar, منات) was a type of artefact closely associated with the goddess Hathor. Operation The menat was held in the hand by its counterpoise and used as a rattle by Hathor's priestesses. It ...
maar A maar is a broad, low-relief volcanic crater caused by a phreatomagmatic eruption (an explosion which occurs when groundwater comes into contact with hot lava or magma). A maar characteristically fills with water to form a relatively shallow ...
deposit in France bears similarities to both Messelasturidae and Halcyornithidae.


References

{{Taxonbar, from=Q2558536 Parrots Eocene birds Prehistoric birds of Europe Prehistoric bird families