Mesophoyx Intermedia
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The intermediate egret, median egret, smaller egret or medium egret (''Ardea intermedia'') is a medium-sized
heron The herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 72 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genera ''Botaurus'' and ''Ixobrychus ...
. Some taxonomists put the species in the genus ''
Egretta ''Egretta'' is a genus of medium-sized herons, mostly breeding in warmer climates. Representatives of this genus are found in most of the world, and the little egret, as well as being widespread throughout much of the Old World, has now started ...
'' or ''Mesophoyx''. It is a resident breeder in southern and eastern Asia.


Taxonomy

Some authorities classify the intermediate egret in its own
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus, ''Mesophoyx'', while others place it with the smaller egrets in ''
Egretta ''Egretta'' is a genus of medium-sized herons, mostly breeding in warmer climates. Representatives of this genus are found in most of the world, and the little egret, as well as being widespread throughout much of the Old World, has now started ...
''. There were three recognised subspecies, and these are sometimes raised in to species: * '' A. i. brachyrhyncha'' Brehm, 1854 "yellow-billed egret" - sub-Saharan Africa * ''A.i. intermedia'' Wagler, 1827 "intermediate egret" - Asia from the Russian Far East to Japan to India and the
Greater Sundas The Greater Sunda Islands (Indonesian and Malay: ''Kepulauan Sunda Besar'') are four tropical islands situated within Indonesian Archipelago, in the Pacific Ocean. The islands, Borneo, Java, Sulawesi and Sumatra, are internationally recognised fo ...
* '' A. i. plumifera''
Gould Gould may refer to: People * Gould (name), a surname Places United States * Gould, Arkansas, a city * Gould, Colorado, an unincorporated community * Gould, Ohio, an unincorporated community * Gould, Oklahoma, a town * Gould, West Virginia, a ...
, 1848
"plumed egret" - eastern Indonesia,
New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...
and Australia. ''A.(i.) intermedia'' differs from ''A.(i.) brachyrhyncha'' and ''A. (i.) plumifera'' by having a black bill when in breeding plumage, while ''A.(i.) plumifera'' has a yellow-and-pink bill and ''A. (i.) brachyrhyncha'' has much yellower lores and face. A further difference between this species and the "yellow-billed" species is that the intermediate egret has black at the top of the legs compared to reddish in the yellow-billed egret. The split of intermediate egret into 3 species was accepted by the IOC on 26 September 2023, this form was given the common name medium egret by the IOC.


Description

This species, as its scientific name implies, is intermediate in size between the
great egret The great egret (''Ardea alba''), also known as the common egret, large egret, or (in the Old World) great white egret or great white heron is a large, widely distributed egret. The four subspecies are found in Asia, Africa, the Americas, and ...
and smaller white egrets like the little egret and
cattle egret The cattle egret (''Bubulcus ibis'') is a cosmopolitan species of heron (family Ardeidae) found in the tropics, subtropics, and warm-temperate zones. It is the only member of the monotypic genus ''Bubulcus'', although some authorities regard it ...
, though nearer to little than great. It is about long with a wingspan and weighs c. , with all-white plumage, generally dark legs and a thickish yellow bill. Breeding birds may have a reddish or black bill, greenish yellow
gape The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food, ...
skin, loose filamentous plumes on their breast and back, and dull yellow or pink on their upper legs (regional variations). The sexes are similar.


Differences from other egrets

The non-breeding colours are similar, but the intermediate is smaller, with neck length a little less than body length, a slightly domed head, and a shorter, thicker bill. The
great egret The great egret (''Ardea alba''), also known as the common egret, large egret, or (in the Old World) great white egret or great white heron is a large, widely distributed egret. The four subspecies are found in Asia, Africa, the Americas, and ...
has a noticeable kink near the middle of its neck, and the top of its longer bill nearly aligns with the flat top of its head. Close up, great egret's gape line extends behind the eye, while the intermediate's is less pointed and ends below the eye. The intermediate tends to stalk upright with neck extended forward. The great is more patient, often adopting a sideways-leaning "one-eyed" stance. Little egrets have yellow-soled feet and black bills. They often run after fish in shallow water. Breeding birds have long nuptial plumes on the back of their heads.


Behaviour

The intermediate egret stalks its prey methodically in shallow coastal or fresh water, including flooded fields. It eats fish, frogs, crustaceans and insects. It often nests in
colonies In modern parlance, a colony is a territory subject to a form of foreign rule. Though dominated by the foreign colonizers, colonies remain separate from the administration of the original country of the colonizers, the '' metropolitan state'' ...
with other herons, usually on platforms of sticks in trees or shrubs. The typical clutch size is 2 or 3 eggs but there can be as many as 6 eggs in a clutch, the colour of the eggs is pale green, with a smooth, slightly pitted shell. Both parents incubate the eggs and they hatch after between 24 and 27 days. The eggs hatch asynchronously, after hatching the adults brood the semialtricial young for 12 days, defending the nests from aerial predators during both incubation and brooding, they crouch over the nest and raise their plumes and point their bill towards the threat. The parents regurgitate the food to feed the young, initially onto the floor of the nest but later the chick takes the food from the parent's mouth. There may be competition for food within the brood. Pinfeathers may appear on the chicks as early as 4 days old and the young are able to leave the nest at 24 days old, although they return to be fed. Fledging occurs at around 40 days old and they leave the colony after 70 days. In Africa and Australia they are reported to be quite successful breeders with 96% and 88% of nests fledging at least a single chick. Intermediate egret populations are more successful in wet years than in dry years.


Gallery

File:Inter Egret Descending Tree Kabini Apr22 D72 23600.jpg, Descending to perch,
Kabini River The Kabini River is one of the major tributaries of the river Cauvery in southern India. It originates near Kavilumpara in Kozhikode district of Kerala state by the confluence of the Panamaram River and the Mananthavady River. It flows eastw ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
File:Intermediate Egret day95.ogg, Feeding on mice, SE Queensland, Australia File:Egret's tongue.JPG, Egret showing its tongue File:Photo showing the size difference in Great, Intermediate and Little Egrets - Taken From Dhikala, Jim Corbett TR.jpg, Photo showing the size difference in great, intermediate and little egrets, India File:Intermediate Egret Feeding.2.jpg, In
Northern Territory The Northern Territory (commonly abbreviated as NT; formally the Northern Territory of Australia) is an states and territories of Australia, Australian territory in the central and central northern regions of Australia. The Northern Territory ...
, Australia


References

* ''Birds of The Gambia'' by Barlow, Wacher and Disley, * ''Birds of India'' by Grimmett, Inskipp and Inskipp,


External links


Species text in ''The Atlas of Southern African Birds''
{{Taxonbar, from=Q27074615, from2=Q218821
intermediate egret The intermediate egret, median egret, smaller egret, or yellow-billed egret (''Ardea intermedia'') is a medium-sized heron. Some taxonomists put the species in the genus ''Egretta'' or ''Mesophoyx''. It is a resident breeder from east Africa a ...
Birds of Japan Birds of East Asia Birds of South Asia Birds of Southeast Asia Birds of Sub-Saharan Africa
intermediate egret The intermediate egret, median egret, smaller egret, or yellow-billed egret (''Ardea intermedia'') is a medium-sized heron. Some taxonomists put the species in the genus ''Egretta'' or ''Mesophoyx''. It is a resident breeder from east Africa a ...
intermediate egret The intermediate egret, median egret, smaller egret, or yellow-billed egret (''Ardea intermedia'') is a medium-sized heron. Some taxonomists put the species in the genus ''Egretta'' or ''Mesophoyx''. It is a resident breeder from east Africa a ...
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