Megabalanus Rosa
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''Megabalanus'' is a genus of
barnacle A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in eros ...
s in the family
Balanidae The Balanidae comprise a family of barnacles of the order Balanomorpha. As a result of research published in 2021 by Chan et al., the members of the family Archaeobalanidae were merged with this family. Genera These genera belong to the family B ...
. Members of the genus grow to in length and inhabit the lower
intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species o ...
.


Morphology

''Megabalanus'' is an
acorn barnacle Acorn barnacle and acorn shell are Common name, vernacular names for certain types of stalkless barnacles, generally excluding goose barnacles, stalked or gooseneck barnacles. As adults they are typically cone-shaped, symmetrical, and attached to r ...
, a sessile crustacean that secretes a calcium carbonate shell consisting of five plates. It reaches up to in height .


Ecology

Like other barnacles, they are
suspension feeder Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...
s, grabbing particulate matter from the water column overhead. ''M. tintinnabulum'' has a worldwide distribution and is commonly found on wave-beaten shores, in the lower intertidal zone. They tend to grow in clusters of around 12 individuals. Other species are more select, such as ''M. californicus'' of tidal habitats in California and the greater North American Pacific Coast. While many barnacles deal with competition for space by organisms such as
limpet Limpets are a group of aquatic snails that exhibit a conical shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended indep ...
s and
mussel Mussel () is the common name used for members of several families of bivalve molluscs, from saltwater and Freshwater bivalve, freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other ...
s by growing many organisms close together in colonies, ''Megabalanus'' responds by rapidly growing to a very large size. Their large size also helps reduce predation, although it makes them large enough to be harvested for human consumption.


Geological utility

Fossilised specimens of ''Megabalanus'' have been found dating back to the Miocene. Fossils bearing a close resemblance to ''M. tintinnabulum'' are preserved in large numbers in the Tabernas Basin of Spain. A case study of this area showed that the state of preservation of the organisms makes possible estimating the distance they were transported ''post mortem'' . Since the species today lives in the intertidal zone, this allows an estimate to be made of the distance from the shoreline, and by implication, the water depth. These observations are backed up by the appearance of the deep-water trace fossil '' Zoophycum'' in the predicted deepest waters, suggesting that barnacles can be a good proxy for water depth.


Species

The World Register of Marine Species includes these species in the genus: * ''
Megabalanus ajax ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Darwin, 1854) * ''
Megabalanus antillensis ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Pilsbry, 1916) * ''
Megabalanus azoricus ''Megabalanus azoricus'', the Azorean barnacle, is a species of large barnacle in the family Balanidae. It is endemic to Macaronesia: the Azores, Madeira, the Canary Islands, and Cape Verde. It is the largest and third most common shallow water b ...
'' (Pilsbry, 1916) (Azorean barnacle) * ''
Megabalanus californicus ''Megabalanus californicus'', the California barnacle, is a species of large barnacle in the family Balanidae. It is native to rocky coasts in the Eastern Pacific Ocean from North California to the Gulf of California. Description ''Megabalanus c ...
'' (Pilsbry, 1916) (California barnacle) * ''
Megabalanus clippertonensis ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' Zullo, 1969 * ''
Megabalanus coccopoma ''Megabalanus coccopoma'', the titan acorn barnacle, is a tropical species of barnacle first described by Charles Darwin in 1854. Its native range is the Pacific coasts of South and Central America but it is extending its range to other parts of ...
'' (Darwin, 1854) (titan acorn barnacle) * ''
Megabalanus costatus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Hoek, 1913) * ''
Megabalanus crispatus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Schröter) Darwin, 1854 * ''
Megabalanus dolfusii ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (de Alessandri, 1907) * ''
Megabalanus dorbignii ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (chenu, 1843) * ''
Megabalanus galapaganus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Pilsbry, 1916) * ''
Megabalanus giganteum ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Kolosváry, 1949) * '' Megabalanus honti'' (Kolosváry, 1950) * ''
Megabalanus hungaricus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Kolosváry, 1941) * ''
Megabalanus javanicus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Whithers, 1923) * ''
Megabalanus leganyii ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Kolosváry, 1950) * ''
Megabalanus linzei ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Foster, 1978) * ''
Megabalanus multiseptatus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Ross, 1964) * '' Megabalanus occator'' (Darwin, 1854) * ''
Megabalanus peninsularis ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbo ...
'' (Pilsbry, 1916) * ''
Megabalanus plicatus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Hoek, 1913) * '' Megabalanus rosa'' Pilsbry, 1916 (acorn barnacle) * ''
Megabalanus seguenzai ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (de Alessandri, 1895) * ''
Megabalanus spinosus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Gmelin, 1791) * ''
Megabalanus stultus ''Megabalanus stultus'' is a species of barnacle first described by Charles Darwin in 1854. It lives on fire corals of the genus ''Millepora'' in the Atlantic Ocean from Florida to southern Brazil. Ecology ''Megabalanus stultus'' is one of thre ...
'' (Darwin, 1854) * '' Megabalanus tanagrae'' (Pilsbry, 1928) * ''
Megabalanus tintinnabulum ''Megabalanus tintinnabulum'' is a species of large barnacle in the family Balanidae. It is the type species of the genus. The specific name comes from the Latin ' meaning a handbell and probably refers to the fact that small groups of barnacles ...
'' ( Linnaeus,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of Systema Naturae, 10th edition of ''Systema Naturae'', the sta ...
)
(titan acorn barnacle) * ''
Megabalanus transsylvanicus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Kolosváry, 1950) * '' Megabalanus transversostriatus'' (Beurlen, 1958) * '' Megabalanus tulipiformis'' (Ellis, 1758) * '' Megabalanus validus'' Darwin, 1854 * ''
Megabalanus venezuelensis ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbo ...
'' (Weisbord, 1966) * ''
Megabalanus vesiculosus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Darwin, 1854) * ''
Megabalanus vinaceus ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Darwin, 1854) * ''
Megabalanus volcano ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' (Pilsbry, 1916) * ''
Megabalanus xishaensis ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbo ...
'' (Ren & Liu, 1978) * ''
Megabalanus zebra ''Megabalanus'' is a genus of barnacles in the family Balanidae. Members of the genus grow to in length and inhabit the lower intertidal zone. Morphology ''Megabalanus'' is an acorn barnacle, a sessile crustacean that secretes a calcium carbon ...
'' Russell et al., 2003


References

{{Taxonbar, from=Q3282981 Barnacles Extant Miocene first appearances