Mecodema Alternans
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''Mecodema'' is a genus of large flightless ground beetle (Carabidae) endemic to New Zealand. The genus is very diverse in comparison to the other three New Zealand genera ('' Diglymma'', '' Oregus'', '' Orthoglymma'') within the subtribe Nothobroscina (tribe Broscini). ''Mecodema'' is geographically widespread across both the North and
South Islands The Kingdom of the Isles consisted of the Isle of Man, the Hebrides and the islands of the Firth of Clyde from the 9th to the 13th centuries AD. The islands were known to the Norse as the , or "Southern Isles" as distinct from the or Nort ...
, as well as numerous offshore islands, including the Three Kings Is., Poor Knights Is., Aotea ( Great Barrier Is.) and Hauturu (Little Barrier Is.), Kapiti Is., Stephens Is., Stewart Is., Chatham Is., Snares Is.


Genus description

Head, frons and vertex often rugose and / or punctate, microsculpture or macrosculpture absent in some species groups (e.g., ''curvidens'' species); eyes prominent, almost hemispherical; a single supraorbital puncture on each side, bearing more than one setae; setose punctures of vertex absent; mandibles large, uniformly narrow to the pointed apex, sharply curved in the apical third, mandibles with a number of dorsal grooves and without a seta in the scrobe; mentum with median process, which is notched to strongly indentate (bifid); usually with 2 setiferous punctures (may be absent) below the process and generally with 4–8 setose (setiferous) punctures along the submentum sclerite; labial palpi with setae on the penultimate segment; terminal segments of all palpi slightly compressed, sub-cylindrical, truncate; ligula chitinous with a median carina, 2 apical setae present; the membranous paraglossae extended well beyond the apex of the ligula; stipes with 2–3 basal setae; antennomeres I–II glabrous, III–V with apical ring, VI–XI setose throughout; antennomeres I–III subcylindrical, IV–VII subglobose and slightly compressed. Prothoracic carina (pronotal lateral margin) distinct with an asymmetrical number of setae along each side, the last never at the posterior lateral sinuation (posterior angle); midline (median groove) and pronotal foveae (laterobasal depressions) always present; pronotal disc often with shallow depressions on each side of apical half. Elytra always striate; fused together along the suture; hind wings absent; a row of setose punctures on the 7th stria and another row (umbilical series) near the lateral carina (elytral margin). Legs slender, the protibia with a pointed apical prolongation on the outer side, greatly distally expanded to distally expanded and shovel-like; male and female tarsi identical, the basal 4 segments of the anterior tarsi are as broad as they are long, usually slightly asymmetrical. Type species: '' Mecodema sculpturatum''
Blanchard Blanchard is a French family name. It is also used as a given name. It derives from the Old French word ''blanchart'' which meant "whitish, bordering upon white". It is also an obsolete term for a white horse. Geographical distribution As of 2014, ...
1853.


Species and subspecies (102)

* '' Mecodema aberrans'' (Putzeys, 1868) * '' Mecodema allani'' Fairburn, 1945 * '' Mecodema alternans'' Laporte de Castelnau, 1867 ** ''Mecodema alternans hudsoni''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1909 * ''
Mecodema angustulum ''Mecodema'' is a genus of large flightless ground beetle (Carabidae) endemic to New Zealand. The genus is very diverse in comparison to the other three New Zealand genera ('' Diglymma'', '' Oregus'', '' Orthoglymma'') within the subtribe Noth ...
''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1914 * '' Mecodema antarcticum'' (Laporte de Castelnau, 1867) * '' Mecodema aoteanoho'' Seldon & Leschen, 2011 * '' Mecodema argentum'' Seldon & Buckley, 2019 * '' Mecodema atrox'' Britton, 1949 * '' Mecodema atuanui'' Seldon & Buckley, 2019 * '' Mecodema brittoni'' Townsend, 1965 * '' Mecodema bullatum'' Lewis, 1902 * '' Mecodema chaiup'' Seldon, 2015 * '' Mecodema chiltoni''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1917 * '' Mecodema constrictum'' Broun, 1881 * '' Mecodema costellum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1903 ** ''Mecodema costellum gordonense''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1917 ** ''Mecodema costellum lewisi''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1908 ** ''Mecodema costellum obesum'' Townsend, 1965 * '' Mecodema costipenne''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1914 * '' Mecodema crenaticolle'' Redtenbacher, 1868 * '' Mecodema crenicolle'' Castelnau, 1867 * ''
Mecodema curvidens ''Mecodema curvidens'' Broun is a medium-bodied ground beetle that is geographically widespread throughout the central areas of the North Island The North Island, also officially named Te Ika-a-Māui, is one of the two main islands of New ...
'' (
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1915) * '' Mecodema ducale'' Sharp, 1886 * '' Mecodema dunense'' Townsend, 1965 * '' Mecodema dunnorum'' Seldon & Buckley, 2019 * '' Mecodema dux'' Britton, 1949 * '' Mecodema elongatum'' Castelnau, 1867 * '' Mecodema femorale''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1921 * '' Mecodema florae'' Britton, 1949 * '' Mecodema fulgidum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1881 * '' Mecodema genesispotini'' Seldon & Buckley, 2019 * '' Mecodema godzilla'' Seldon & Buckley, 2019 * '' Mecodema gourlayi'' Britton, 1949 * '' Mecodema haunoho'' Seldon & Leschen, 2011 * '' Mecodema hector'' Britton, 1949 * '' Mecodema howitti'' Castelnau, 1867 * '' Mecodema huttense''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1915 * '' Mecodema impressum'' Castelnau, 1867 * '' Mecodema infimate'' Lewis, 1902 * '' Mecodema integratum'' Townsend, 1965 * ''
Mecodema jacinda ''Mecodema jacinda'' is a large-bodied ground beetle endemic to Maungatautari Sanctuary, Waikato, New Zealand. It is the largest species of ''Mecodema'' found on Maungatautari, which also has the medium-sized '' M. curvidens'' inhabiting the fo ...
'' Seldon & Buckley, 2019 * '' Mecodema kipjac'' Seldon & Buckley, 2019 * '' Mecodema kokoroiho'' Seldon & Buckley, 2019 * '' Mecodema kokoromatua'' Seldon, Leschen & Liebherr, 2012 * '' Mecodema laeviceps''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1904 * '' Mecodema laterale''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1917 * '' Mecodema litoreum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1886 * '' Mecodema longicolle''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1923 * '' Mecodema lucidum'' Castelnau, 1867 * '' Mecodema manaia'' Seldon & Leschen, 2011 * '' Mecodema metallicum'' Sharp, 1886 * '' Mecodema minax'' Britton, 1949 * '' Mecodema mohi'' Seldon & Buckley, 2019 * '' Mecodema moniliferum'' (Bates, 1867) * '' Mecodema morio'' ( Castelnau, 1867) * '' Mecodema ngaiatonga'' Seldon & Buckley, 2019 * '' Mecodema ngaitahuhu'' Seldon & Buckley, 2019 * '' Mecodema nitidum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1903 * '' Mecodema oblongum'' (
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1882) * '' Mecodema oconnori''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1912 * '' Mecodema oregoides'' (
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1894) * '' Mecodema papake'' Seldon & Buckley, 2019 * '' Mecodema parataiko'' Seldon & Leschen, 2011 * '' Mecodema pavidum'' Townsend, 1965 * '' Mecodema perexiguus'' Seldon & Buckley, 2019 * '' Mecodema persculptum'' Broun, 1915 * '' Mecodema pluto'' Britton, 1949 * '' Mecodema politanum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1917 * '' Mecodema ponaiti'' Seldon & Leschen, 2011 * '' Mecodema proximum'' Britton, 1949 * '' Mecodema puiakium'' Johns & Ewers in Johns, 2007 * '' Mecodema pulchellum'' Townsend, 1965 * '' Mecodema punctatum'' (Laporte de Castelnau, 1867) * '' Mecodema punctellum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1921 * '' Mecodema puncticolle''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1915 Mecodema sculpturatum'' Blanchard, 1843">='' Mecodema sculpturatum'' Blanchard, 1843* '' Mecodema quoinense''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1912 * '' Mecodema rectolineatum'' Laporte de Castelnau, 1867 * '' Mecodema regulus'' Britton, 1964 * '' Mecodema rex'' Britton, 1949 * '' Mecodema rugiceps'' Sharp, 1886 ** ''Mecodema rugiceps anomalum'' Townsend, 1965 * '' Mecodema rusticulus'' Seldon & Buckley, 2019 * '' Mecodema scitulum'' Broun, 1894 * '' Mecodema sculpturatum'' Blanchard, 1843 * '' Mecodema simplex'' Castelnau, 1867 * '' Mecodema spiniferum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1880 * '' Mecodema striatum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1904 * '' Mecodema strictum'' Britton, 1949 * '' Mecodema temata'' Seldon & Buckley, 2019 * '' Mecodema tenaki'' Seldon & Leschen, 2011 * '' Mecodema teparawhau'' Seldon & Buckley, 2019 * '' Mecodema teroroa'' Seldon & Buckley, 2019 * '' Mecodema tewhara'' Seldon & Buckley, 2019 * '' Mecodema tibiale'' (Laporte de Castelnau, 1867) * '' Mecodema tuhoe'' Seldon & Buckley, 2019 * '' Mecodema undecimus'' Seldon & Buckley, 2019 * '' Mecodema validum''
Broun Broun is a surname. It is the Middle English and Scots spelling of Brown. Notable people with the surname include: *Agnes Broun (1732–1820), mother of Scottish poet Robert Burns *Alex Broun (born 1965), Australian playwright and screenwriter *Da ...
, 1923 * '' Mecodema wharekahika'' Seldon & Buckley, 2019 * '' Mecodema xylanthrax'' Seldon & Buckley, 2019 * '' Mecodema yconomus'' Seldon & Buckley, 2019 * '' Mecodema zonula'' Seldon & Buckley, 2019


References


Further reading

* Seldon, D. S. (2015). A unique species of ''Mecodema'' (Carabidae: Broscini) from the Hawke's Bay region, New Zealand, with implications for North–South Island zoogeography. ''New Zealand Entomologist'', ''38''(1), 28-35. * Seldon, D. S., & Leschen, R. A. (2011). Revision of the ''Mecodema curvidens'' species group (Coleoptera: Carabidae: Broscini). ''Zootaxa'', ''2829''(1), 1-45. * Seldon, D. S., Leschen, R. A. B., & Liebherr, J. K. (2012). A new species of ''Mecodema'' (Carabidae: Broscini) from Northland, New Zealand, with notes on a newly observed structure within the female genitalia. ''New Zealand Entomologist'', ''35''(1), 39-50. * Townsend, J. I. (1965). Notes on the genus ''Mecodema'' (Coleoptera: Carabidae) with descriptions of new species from the South Island of New Zealand. ''New Zealand Journal of Science'', ''8''(3), 301-318. * Townsend, J. I. (1971). Entomology of the Aucklands and other islands south of New Zealand: Coleoptera: Carabidae: Broscini. ''Pacific Insects Monograph'', ''27'', 173-184.
NZ Entomologist
*


Landcare Research
{{Taxonbar, from=Q1382067 Carabidae genera