''Gymnosporia heterophylla'', the common spike-thorn, is a small, hardy, deciduous

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

n tree up to 5m tall, occurring in rocky places with a wide distribution from

Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...

, the

Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...

and the Congo, south to the

Cape Province The Province of the Cape of Good Hope ( af, Provinsie Kaap die Goeie Hoop), commonly referred to as the Cape Province ( af, Kaapprovinsie) and colloquially as The Cape ( af, Die Kaap), was a province in the Union of South Africa and subsequen ...

and west to

Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...

and

Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...

, as well as the neighbouring islands of

Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...

and

Saint Helena Saint Helena () is a British overseas territory located in the South Atlantic Ocean. It is a remote volcanic tropical island west of the coast of south-western Africa, and east of Rio de Janeiro in South America. It is one of three constitu ...

, with a closely related species from

Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label=Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It incl ...

. It has a straggly, but rigid habit and is armed with sharp straight

thorns Thorn(s) or The Thorn(s) may refer to: Botany * Thorns, spines, and prickles, sharp structures on plants * ''Crataegus monogyna'', or common hawthorn, a plant species Comics and literature * Rose and Thorn, the two personalities of two DC Com ...

up to 100mm long, which are modified branches. Bark on the mature trunk is grey-brown and deeply fissured. The tree is

dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...

, and clusters of white flowers are produced in profusion in spring and are borne on thicker twigs and branches. The flowers have a fetid, faecal smell and attract large numbers of pollinating insects, particularly carrion-loving flies such as members of the family

Calliphoridae The Calliphoridae (commonly known as blow flies, blow-flies, carrion flies, bluebottles, greenbottles, or cluster flies) are a family of insects in the order Diptera, with almost 1,900 known species. The maggot larvae, often used as fishing ba ...

. The tree has a close relationship with a number of

cicada The cicadas () are a superfamily, the Cicadoidea, of insects in the order Hemiptera (true bugs). They are in the suborder Auchenorrhyncha, along with smaller jumping bugs such as leafhoppers and froghoppers. The superfamily is divided into two ...

species, such as '' Platypleura divisa'', '' P. mijburghi'' and '' P. maytenophila''. The specific name "heterophylla" means "variable leaves" and if the list of past synonyms is examined, it affords an interesting insight into the minds of botanists since 1753 when Linnaeus decided that the leaves reminded him of

boxwood ''Buxus'' is a genus of about seventy species in the family Buxaceae. Common names include box or boxwood. The boxes are native to western and southern Europe, southwest, southern and eastern Asia, Africa, Madagascar, northernmost South ...

and named it ''Celastrus buxifolius''. Since then it has been named ''ellipticus, heterophyllus, spathephyllus, empleurifolius, rhombifolius, parvifolius and buxifolioides'' - it would seem that the epithet ''heterophylla'' is appropriate. It makes a very effective, fast-growing security hedge. The wood is hard and durable, suitable for carving, though large pieces are not often available.

Pictures

File:Maytenus heterophylla00.jpg File:Maytenus heterophylla01.jpg

References

''Reinstatement of "Gymnosporia"
*https://plants.jstor.org/compilation/maytenus.heterophylla *

Sources

*Trees and Shrubs of the Witwatersrand - Tree Society (

Witwatersrand University The University of the Witwatersrand, Johannesburg (), is a multi-campus South African public research university situated in the northern areas of central Johannesburg. It is more commonly known as Wits University or Wits ( or ). The university ...

Press, 1969) *Trees and Shrubs of

Mpumalanga Mpumalanga () is a province of South Africa. The name means "East", or literally "The Place Where the Sun Rises" in the Swazi, Xhosa, Ndebele and Zulu languages. Mpumalanga lies in eastern South Africa, bordering Eswatini and Mozambique. It ...

and

Kruger National Park Kruger National Park is a South African National Park and one of the largest game reserves in Africa. It covers an area of in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends from north to south and from ea ...

- Schmidt et al. (Jacana Press, 2002) heterophylla Trees of Africa Trees of South Africa {{tree-stub