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''Maylandia'' or ''Metriaclima'' is a genus of
haplochromine __NOTOC__ The haplochromine cichlids are a tribe of cichlids in subfamily Pseudocrenilabrinae called Haplochromini. This group includes the type genus (''Haplochromis'') plus a number of closely related genera such as '' Aulonocara'', '' Astat ...
cichlid Cichlids are fish from the family Cichlidae in the order Cichliformes. Cichlids were traditionally classed in a suborder, the Labroidei, along with the wrasses ( Labridae), in the order Perciformes, but molecular studies have contradicted this ...
s endemic to Lake Malawi in
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
. They belong to the
mbuna Mbuna (pronounced Mmm-boo - nah ) is the common name for a large group of African cichlids from Lake Malawi, and are members of the haplochromine family. The name ''mbuna'' means "rockfish" in the language of the Tonga people of Malawi.Loiselle ...
(rock-dwelling) haplochromines. All species in this genus are relatively small fishes, less than in length. Like most Lake Malawi cichlids, exhibit brood care via maternal
mouthbrooding Mouthbrooding, also known as oral incubation and buccal incubation, is the care given by some groups of animals to their offspring by holding them in the mouth of the parent for extended periods of time. Although mouthbrooding is performed by a va ...
. Numerous members of the genus are traded as aquarium fish. They are attractive because they are brightly colored and often very sexually dimorphic; like other
cichlid Cichlids are fish from the family Cichlidae in the order Cichliformes. Cichlids were traditionally classed in a suborder, the Labroidei, along with the wrasses ( Labridae), in the order Perciformes, but molecular studies have contradicted this ...
s they are not suited for beginners and for most companion tanks.


Name

The name ''Maylandia'', honours the cichlid enthusiast and author about aquaria Hans Joachim Mayland, who died in 2004, was proposed as a
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
of ''
Pseudotropheus ''Pseudotropheus'' is a genus of fishes in the family Cichlidae. These mbuna cichlids are endemic to Lake Malawi in Eastern Africa. Taxonomy Like some other large cichlid genera, such as ''Cichlasoma'', a number of related fishes have been recen ...
'' in 1984, naming the long-known but undescribed "
Ice Blue Zebra William's mbuna (''Maylandia greshakei''), also known as ice blue zebra mbuna or ice blue zebra or the Pseudotropheus ice blue among the aquarium enthusiasts, is a species of cichlid fish endemic to Lake Malawi where it is only found at Makok ...
" as the type species. In 1997 Stauffer ''et al''. described the genus ''Metriaclima'', dismissing the pre-existing ''Maylandia'' on the assumption that it lacked a type species and a diagnosis. Two years later Condé and Géry published an analysis and declared ''Metriaclima'' to be a junior synonym of ''Maylandia'', and ''Maylandia'' hence the valid name of the genus, a view accepted by most ichthyologists. A few authors, notably
Ad Konings Adrianus Franciscus Johannes Marinus Maria "Ad" Konings (born 11 January 1956 in Roosendaal, Netherlands) is an ichthyologist originally trained in medicine and biology. Konings is best known for his research on African rift lake cichlids. After ...
, dispute that the original description of ''Maylandia'' is sufficient to establish and maintain the genus. They maintain that "Maylandia" is a ''
nomen nudum In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
'' – literally, a "naked name" that does not validly refer to a group of animals as per the rules of the '' International Code of Zoological Nomenclature''. In the present article, following
FishBase FishBase is a global species database of fish species (specifically finfish). It is the largest and most extensively accessed online database on adult finfish on the web.
, Catalog of Fishes and the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
, the genus name ''Maylandia'' is used.


Species

There are currently 30 recognized species in this genus: * ''
Maylandia aurora ''Maylandia aurora'' is a species of cichlid endemic to Lake Malawi where it is only known from the southern half of Likoma Island where it prefers habitats with rocky substrates close to sand. Males of this species can reach a length of SL ...
'' ( W. E. Burgess, 1976) * ''
Maylandia barlowi ''Maylandia barlowi'' is a species of cichlid endemic to Lake Malawi where it is only known from the area around the Maleri Islands where it prefers areas with soft substrates. This species can reach a length of SL. It is also found in the a ...
'' ( McKaye & Stauffer, 1986) * ''
Maylandia benetos ''Maylandia benetos'' is a species of cichlid endemic to Lake Malawi Lake Malawi, also known as Lake Nyasa in Tanzania and Lago Niassa in Mozambique, is an African Great Lake and the southernmost lake in the East African Rift system, loca ...
'' ( Stauffer, Bowers, Kellogg & McKaye, 1997) * '' Maylandia callainos'' ( Stauffer & Hert, 1992) (Cobalt Zebra, Cobalt Blue Mbuna) * ''
Maylandia chrysomallos ''Maylandia chrysomallos'' is a species of cichlid endemic to Lake Malawi where it is only known from rocky areas around Mumbo Island. This species can reach a length of SL. It is also found in the aquarium trade. References chrysomall ...
'' ( Stauffer, Bowers, Kellogg & McKaye, 1997) * ''
Maylandia cyneusmarginatus ''Maylandia cyneusmarginata'' is a species of fish in the family Cichlidae. It is endemic to Lake Malawi Lake Malawi, also known as Lake Nyasa in Tanzania and Lago Niassa in Mozambique, is an African Great Lake and the southernmost lake in ...
'' ( Stauffer, Bowers, Kellogg & McKaye, 1997) * ''
Maylandia emmiltos ''Maylandia emmiltos'' is a species of cichlid endemic to Lake Malawi where it is only known from the Mpanga Rocks near Chilumba. This species can reach a length of SL. It is also found in the aquarium An aquarium (plural: ''aquariums' ...
'' ( Stauffer, Bowers, Kellogg & McKaye, 1997) * ''
Maylandia estherae ''Maylandia estherae'' (the Red Zebra mbuna, Red Zebra Cichlid , or Esther Grant's Zebra) is a haplochromine cichlid. It is a rock dwelling fish or mbuna from Lake Malawi. This fish, like most cichlids from Lake Malawi, is a mouthbrooder - female ...
'' (
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
, 1995)
(Red Zebra Mbuna, Red Zebra, Esther Grant's Zebra) * '' Maylandia fainzilberi'' Staeck, 1976 * ''
Maylandia flavicauda ''Maylandia'' or ''Metriaclima'' is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. They belong to the mbuna (rock-dwelling) haplochromines. All species in this genus are relatively small fishes, less than in length. Li ...
'' (Li, Konings & Stauffer 2016) Abstract * ''
Maylandia flavifemina ''Maylandia flavifemina'' is a species of haplochromine cichlid which is endemic to Lake Malawi. Description ''Maylandia flavifemina'' has 17–19 spines in its dorsal fin which also have 17–19 soft rays, there are 8-10 spines and 6–9 soft r ...
'' (
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
& Stauffer, 2006)
* ''
Maylandia glaucos ''Maylandia'' or ''Metriaclima'' is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. They belong to the mbuna (rock-dwelling) haplochromines. All species in this genus are relatively small fishes, less than in length. Li ...
'' ( Ciccotta,
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
& Stauffer, 2011)
* ''
Maylandia greshakei William's mbuna (''Maylandia greshakei''), also known as ice blue zebra mbuna or ice blue zebra or the Pseudotropheus ice blue among the aquarium enthusiasts, is a species of cichlid fish endemic to Lake Malawi where it is only found at Makokol ...
'' ( M. K. Meyer & W. Förster, 1984) (William's Mbuna, Ice Blue Zebra Mbuna, Ice Blue Zebra) * ''
Maylandia hajomaylandi ''Maylandia hajomaylandi'' is a species of cichlid endemic to Lake Malawi where it is only known from around Chisumulu Island. This species can reach a length of TL. It can also be found in the aquarium trade. The specific name Specific nam ...
'' ( M. K. Meyer & Schartl, 1984) * ''
Maylandia heteropicta ''Maylandia heteropicta'' is a species of cichlid endemic to Lake Malawi where it is only known from Thumbi West Island. This species can reach a length of SL. It can also be found in the aquarium An aquarium (plural: ''aquariums'' or ...
'' Staeck, 1980 * ''
Maylandia koningsi ''Maylandia'' or ''Metriaclima'' is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. They belong to the mbuna (rock-dwelling) haplochromines. All species in this genus are relatively small fishes, less than in length. Li ...
'' (Stauffer 2018) Abstract * '' Maylandia lanisticola'' ( W. E. Burgess, 1976) * '' Maylandia livingstonii'' (
Boulenger Boulenger is a surname. Notable people with the surname include: * Benjamin Boulenger (born 1990), French footballer * Edward George Boulenger (1888–1946), British zoologist, director of aquarium at London Zoo * George Albert Boulenger (1858–1 ...
, 1899)
* ''
Maylandia lombardoi ''Maylandia lombardoi'', is a long freshwater fish from the family Cichlidae. This species is popular in the aquarium hobby where it is sold under a variety of common names including: lombardoi mbuna, kenyi mbuna or kennyi mbuna or kenyi cichlid ...
'' ( W. E. Burgess, 1977) (Kennyi Mbuna, Kenyi Mbuna, Lombardoi Mbuna) * ''
Maylandia lundoensis ''Maylandia'' or ''Metriaclima'' is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. They belong to the mbuna (rock-dwelling) haplochromines. All species in this genus are relatively small fishes, less than in length. Li ...
'' ( Stauffer, K. E. Black &
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
, 2013)
Stauffer, J.R. Jr., Black, K. & Konings, A.F. (2013)
Descriptions of five new species of ''Metriaclima'' (Teleostei: Cichlidae) from Lake Malawi, Africa.
''Zootaxa, 3647 (1): 101–136.''
* ''
Maylandia mbenjii ''Maylandia mbenjii'' is a species of cichlid endemic to Lake Malawi where it is only known from Mbenji Island. This species can reach a length of SL. It is also found in the aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') ...
'' ( Stauffer, Bowers, Kellogg & McKaye, 1997) * ''
Maylandia midomo ''Maylandia'' or ''Metriaclima'' is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. They belong to the mbuna (rock-dwelling) haplochromines. All species in this genus are relatively small fishes, less than in length. Li ...
'' ( Stauffer, K. E. Black &
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
, 2013)
* '' Maylandia mossambica'' ( Ciccotta,
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
& Stauffer, 2011)
* '' Maylandia nigrodorsalis'' ( Stauffer, K. E. Black &
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
, 2013)
* '' Maylandia nkhunguensis'' ( Ciccotta,
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
& Stauffer, 2011)
* ''
Maylandia pambazuko ''Maylandia'' or ''Metriaclima'' is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. They belong to the mbuna (rock-dwelling) haplochromines. All species in this genus are relatively small fishes, less than in length ...
'' ( Stauffer, K. E. Black &
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
, 2013)
* ''
Maylandia phaeos ''Maylandia phaeos'' is a species of cichlid endemic to Lake Malawi where it is known from Cobue on the Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhik ...
'' ( Stauffer, Bowers, Kellogg & McKaye, 1997) * ''
Maylandia pulpican ''Maylandia'' or ''Metriaclima'' is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. They belong to the mbuna (rock-dwelling) haplochromines. All species in this genus are relatively small fishes, less than in length. Li ...
'' (
Tawil ''Ṭawīl'' ( ar, طويل, literally 'long'), or ''al-Ṭawīl'' (), is a meter used in classical Arabic poetry Arabic poetry ( ar, الشعر العربي ''ash-shi‘ru al-‘Arabīyyu'') is the earliest form of Arabic literature. Present kn ...
, 2002)
* ''
Maylandia pyrsonotos ''Maylandia pyrsonotos'' is a species of cichlid Endemism, endemic to Lake Malawi where it occurs naturally around Nakatenga Island and has been introduced to the waters around Maleri Island. This species can reach a length of fish measurement, ...
'' ( Stauffer, Bowers, Kellogg & McKaye, 1997) * ''
Maylandia sciasma ''Maylandia'' or ''Metriaclima'' is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. They belong to the mbuna (rock-dwelling) haplochromines. All species in this genus are relatively small fishes, less than in length. Li ...
'' ( Ciccotta,
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
& Stauffer, 2011)
* ''
Maylandia tarakiki ''Maylandia'' or ''Metriaclima'' is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. They belong to the mbuna (rock-dwelling) haplochromines. All species in this genus are relatively small fishes, less than in length ...
'' ( Stauffer, K. E. Black &
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
, 2013)
* ''
Maylandia xanstomachus ''Maylandia xanstomachus'' is a species of cichlid endemic to Lake Malawi where it is only known from the Maleri Islands and Kanjedza Island. This species can reach a length of TL. It can also be found in the aquarium An aquarium (plural ...
'' ( Stauffer & Boltz, 1989) * '' Maylandia xanthos'' ( Ciccotta,
Konings Konings is a Dutch surname and may refer to: *Ad Konings (b. 1956), Dutch ichthyologist *Anthony Konings (1821–1884), Dutch Catholic priest and Redemptorist professor *Jean Konings (1886–1974), Belgian sprinter *Giovanni Matteo Konings ...
& Stauffer, 2011)
* '' Maylandia zebra'' (
Boulenger Boulenger is a surname. Notable people with the surname include: * Benjamin Boulenger (born 1990), French footballer * Edward George Boulenger (1888–1946), British zoologist, director of aquarium at London Zoo * George Albert Boulenger (1858–1 ...
, 1899)
(Zebra Mbuna)


Footnotes


References

* (2010)

"''Maylandia'' or ''Metriaclima'' – the case for ''Metriaclima''. ''eggspots'' no. 4: 39-45. * (2007)
New ''Metriaclima'' described
'' Practical Fishkeeping, Pract. Fishkeeping''. Retrieved 2009-OCT-03. * (1999): ''Maylandia'' Meyer et Foerster, 1984, un nom générique disponible (Teleostei, Perciformes, Cichlidae) 'Maylandia'' Meyer and Foerster, 1984, an available generic name ''Revue Française d'Aquariologie et de Herpetologie'' 26:(1-2) 21-22. *
009 009 may refer to: * OO9, gauge model railways * O09, FAA identifier for Round Valley Airport * 0O9, FAA identifier for Ward Field, see List of airports in California * British secret agent 009, see 00 Agent * BA 009, see British Airways Flight 9 * ...
br>''Maylandia'' species
Retrieved 2009-OCT-03. * (2009):
2009 IUCN Red List of Threatened Species
'. Version 2009.1. Retrieved 2009-SEP-20. * (1998): ''Maylandia'' oder ''Metriaclima'' ''Maylandia'' or ''Metriaclima''" ''Die Aquarien- und Terrarienzeitschrift'' 51: 813
n German N, or n, is the fourteenth letter in the Latin alphabet, used in the modern English alphabet, the alphabets of other western European languages and others worldwide. Its name in English is ''en'' (pronounced ), plural ''ens''. History ...
* (2001): ''Malawi Cichlids in their natural habitat'' (3rd ed.) Cichlid Press, USA. * (2005)
''Maylandia'' or ''Metriaclima'', ...again!
Retrieved 2007-MAR-02. * (2006): Revised diagnosis of ''Metriaclima'' (Teleostei: Cichlidae) with description of a new species from Lake Malawi National Park, Africa. ''Ichthyological Exploration of Freshwaters'' 17: 233-246
PDF abstract and first page
* (1984): Un nouveau ''Pseudotropheus'' du lac Malawi avec des remarques sur le complexe ''Pseudotropheus-Melanochromis'' (Pisces, Perciformes, Cichlidae) A new ''Pseudotropheus'' from Lake Malawi and remarks on the ''Pseudotropheus-Melanochromis'' complex" ''Revue Française d'Aquariologie et de Herpetologie'' 10: 107-112
n French N, or n, is the fourteenth letter in the Latin alphabet, used in the modern English alphabet, the alphabets of other western European languages and others worldwide. Its name in English is ''en'' (pronounced ), plural ''ens''. History ...
br>HTML fulltext
of genus description. * (2002)

Version of 2002-JAN-07. Retrieved 2009-OCT-03. * (2003)

Version of 2003-JAN-08. Retrieved 2009-OCT-03. * (2009)

Version of 2009-APR-24. Retrieved 2009-OCT-03. * (2009)

"''Maylandia'' or ''Metriaclima'' – still a matter of debate?. ''eggspots'' no. 2: 5-15. * (1998): Von ''Roloffia roloffi'' und ''Maylandia hajomaylandi'' On ''Roloffia roloffi'' and ''Maylandia hajomaylandi''" ''Die Aquarien- und Terrarienzeitschrift'' 51: 812-813
n German N, or n, is the fourteenth letter in the Latin alphabet, used in the modern English alphabet, the alphabets of other western European languages and others worldwide. Its name in English is ''en'' (pronounced ), plural ''ens''. History ...
* (1997): A revision of the blue-black ''Pseudotropheus zebra'' (Teleostei: Cichlidae) complex from Lake Malawi, Africa, with a description of a new genus and ten new species. ''Proceedings of the Academy of Natural Sciences of Philadelphia'' 148: 189-230
Abstract and first page image
{{Taxonbar, from=Q136396 * Haplochromini Cichlid genera