Max Carl Anton Fürbringer (January 30, 1846 – March 6, 1920) was a German anatomist, known for his anatomical investigations of
vertebrate
Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...
s and especially for his studies in
ornithology
Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...
on avian
morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
*Morphology (archaeology), study of the shapes or forms of artifacts
*Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
and
classification Classification is a process related to categorization, the process in which ideas and objects are recognized, differentiated and understood.
Classification is the grouping of related facts into classes.
It may also refer to:
Business, organizat ...
. He was responsible for the first major phylogenetic ordering of bird groups based on a large scale study on a combination of skeletal, morphological and anatomical characteristics.
Biography
Max was the first son of Hermine and Karl Fürbringer. Although he was born in Wittenberg, his father's government employment involved travel and he was raised by relatives in Gera. Max was taught and influenced by
Karl Theodor Liebe
Karl Theodor Leopold Liebe (11 February 1828 - 5 June 1894) was a German geologist and ornithologist. A pioneer in bird conservation, he established the first bird protection agency in Germany.
Liebe was born in Neustadt an der Orla. His father ...
, a bird enthusiast who kept many kinds of birds. Max spent his early childhood collecting coins, butterflies, bees, mosses and ferns. Max's brother Paul would later become an eminent physicist. Max studied at the universities of
Berlin
Berlin ( , ) is the capital and largest city of Germany by both area and population. Its 3.7 million inhabitants make it the European Union's most populous city, according to population within city limits. One of Germany's sixteen constitue ...
and
Jena
Jena () is a German city and the second largest city in Thuringia. Together with the nearby cities of Erfurt and Weimar, it forms the central metropolitan area of Thuringia with approximately 500,000 inhabitants, while the city itself has a popu ...
, where he was a student of Karl Gegenbaur. Fürbringer would be one of Gegenbaur's most ardent disciples and supporters. One of Fürbringer teachers was
Ernst Haeckel
Ernst Heinrich Philipp August Haeckel (; 16 February 1834 – 9 August 1919) was a German zoologist, naturalist, eugenicist, philosopher, physician, professor, marine biologist and artist. He discovered, described and named thousands of new sp ...
about whom he wrote glowingly:
"''He stepped into the auditorium, not with the measured step of the professor, but with the triumphant charging-along of an Apollonian youth, hurrying toward the cathedral, a tall, slender, impressive form; ... golden, flying locks, large, blue, flashing eyes—probably the most beautiful man that I had ever seen, and it seemed to me as if the room, which had already been bright, became noticeably brighter...''"
He obtained his doctorate with a thesis on the muscles and bones of
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s. He later worked as a prosector under Gegenbaur at the
University of Heidelberg
}
Heidelberg University, officially the Ruprecht Karl University of Heidelberg, (german: Ruprecht-Karls-Universität Heidelberg; la, Universitas Ruperto Carola Heidelbergensis) is a public research university in Heidelberg, Baden-Württemberg, ...
, followed by professorships in
Amsterdam
Amsterdam ( , , , lit. ''The Dam on the River Amstel'') is the Capital of the Netherlands, capital and Municipalities of the Netherlands, most populous city of the Netherlands, with The Hague being the seat of government. It has a population ...
,
Jena
Jena () is a German city and the second largest city in Thuringia. Together with the nearby cities of Erfurt and Weimar, it forms the central metropolitan area of Thuringia with approximately 500,000 inhabitants, while the city itself has a popu ...
(from 1888) and
Heidelberg
Heidelberg (; Palatine German language, Palatine German: ''Heidlberg'') is a city in the States of Germany, German state of Baden-Württemberg, situated on the river Neckar in south-west Germany. As of the 2016 census, its population was 159,914 ...
(from 1901). In 1901 he replaced Gegenbaur at Heidelberg as professor and stayed on until 1912 when his student (and later son-in-law) Hermann Braus took over. He then served as an ''außerordentliche'' (=extraordinary) professor at the University of Marburg.
In 1888 Fürbringer published a monumental two volume work on the systematics of birds groups. The first volume was on the forelimbs and shoulder girdle of vertebrates while the second examined the characters and systematics of bird groups. One of the ideas that he used was that the connection of the nerve to a muscle tends to be conserved with evolution. He made use of 51 anatomical and morphological characters and compared every pair of bird group and deduced an evolutionary tree of all the extant and extinct bird groups. His evolutionary tree branched in three dimensions which he represented by means of side and cross-sectional views. This representation was probably influenced by the work of Adolf Engler on plants. This was a seminal work in bird systematics which underwent modifications by
Hans Gadow
Hans Friedrich Gadow (8 March 1855 – 16 May 1928) was a German-born ornithologist who worked in Britain. His work on the classification of birds based on anatomical and morphological characters was influential and made use of by Alexander Wetmore ...
and others to become a definitive basis for subsequent work on the avian evolutionary tree well until the arrival of molecular techniques.
Fuerbringer 1888 tree side view 1.jpg, Front view of tree
Fuerbringer 1888 tree side view 2.jpg, Rear view of tree
Horizontal_Planimetric_I.jpg, Cross section, lower
Horizontal_Planimetric_II.jpg, Cross section, middle
Horizontal_Planimetric_III.jpg, Cross section, upper
As a defender of Gegenbaur, Fürbringer went against
Carl Rabl
Carl Rabl (2 May 1853 in Wels, Austria – 24 December 1917 in Leipzig, Germany
Carl Rabl at
archipterygium or how the fin and the limb were related based on conflicting evidence from anatomy and embryology descended into polemical attacks on each other.
Selected writings
* Die Knochen und Muskeln der Extremitäten bei den schlangenähnlichen Saurien ' ("The bones and muscles of the extremities in the snake-like lizards"), 1870.
* ''Zur Entwickelung der Amphibienniere'' ("The development of amphibian kidneys"), 1877.
* ''Vergleichende Anatomie und Entwicklungsgeschichte der Excretionsorgane der Vertebraten'' ("
Comparative anatomy
Comparative anatomy is the study of similarities and differences in the anatomy of different species. It is closely related to evolutionary biology and phylogeny (the evolution of species).
The science began in the classical era, continuing in t ...
and history of development of the excretion organs of the vertebrates"), 1878.
* Untersuchungen zur Morphologie und Systematik der Vögel ' ("Researches on the morphology and systematics of the bird"), 1888. (in two volumes)
* ''Über die spino-occipitalen Nerven der Salachier und Holocephalen und ihre vergleichende Morphologie'', 1897.
* Beitrag zur Systematik und Genealogie der Reptilien ' ("Contribution to the systematics and genealogy of the reptiles"), 1900.
* ''Zur vergleichenden Anatomie des Brustschulterapparates und der Schultermuskeln'' ("Comparative anatomy of the breast-shoulder apparatus and the shoulder muscles") volumes 4 an 5 1900–1902.
* ''Morphologische Streitfragen'' ("Controversial questions in morphology"), 1902.
* ''Abstammung der Säugetiere'' ("Mammal ancestry"), 1905.