Mary Snow
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''for the composer Mary Snow, see Mary McCarty Snow Christine Mary Snow (Pilkington) (1902–1978) was an Oxford botanist who contributed to the study of
geotropism Gravitropism (also known as geotropism) is a coordinated process of differential growth by a plant in response to gravity pulling on it. It also occurs in fungi. Gravity can be either "artificial gravity" or natural gravity. It is a general featu ...
and
phyllotaxis In botany, phyllotaxis () or phyllotaxy is the arrangement of leaves on a plant stem. Phyllotactic spirals form a distinctive class of patterns in nature. Leaf arrangement The basic arrangements of leaves on a stem are opposite and alterna ...
. She is known for her co-development, with her husband Robin Snow, of the "Snow and Snow Rule": a new
primordium A primordium (; plural: primordia; synonym: anlage) in embryology, is an organ or tissue in its earliest recognizable stage of development. Cells of the primordium are called primordial cells. A primordium is the simplest set of cells capable of ...
(such as a leaf) appears in a plant as soon as and in the place where it has enough space to do so. Mary Pilkington was the daughter of wealthy glass manufacturer Alfred 'Cecil' Pilkington (1875–1966). She was born in
Rainhill Rainhill is a village and civil parish within the Metropolitan Borough of St Helens, in Merseyside, England. The population of the civil parish taken at the 2011 census was 10,853. Historically part of Lancashire, Rainhill was formerly a townsh ...
,
Lancashire Lancashire ( , ; abbreviated Lancs) is the name of a historic county, ceremonial county, and non-metropolitan county in North West England. The boundaries of these three areas differ significantly. The non-metropolitan county of Lancash ...
, on 1 Aug 1902, and became an
exhibitioner An exhibition is a type of scholarship award or bursary. United Kingdom and Ireland At the universities of Dublin, Oxford, Cambridge and Sheffield, at some public schools, and various other UK educational establishments, an exhibition is a sma ...
of St Hugh's College, Oxford in 1922. She graduated with
first-class honours The British undergraduate degree classification system is a grading structure for undergraduate degrees or bachelor's degrees and integrated master's degrees in the United Kingdom. The system has been applied (sometimes with significant variati ...
in botany in 1926 and was a research student at St Hugh's in 1925. In 1926 she became the first research student of George Robert Snow (1897–1969), a botanist and fellow of Magdalen College, Oxford, and was awarded the BSc, a research degree, in 1929. (Known by his nickname Robin, he published under the names R. or Robert Snow.) She was elected as a research fellow of
Somerville College Somerville College, a constituent college of the University of Oxford in England, was founded in 1879 as Somerville Hall, one of its first two women's colleges. Among its alumnae have been Margaret Thatcher, Indira Gandhi, Dorothy Hodgkin, ...
but was unable to take up the position because of her planned marriage to her supervisor Robin Snow in 1930. She continued to teach at Somerville and was later made an honorary research fellow both there and at St Hughs. From 1947 to 1958 she was curator of the botanic gardens. After a trip to Jamaica in 1924, Robin Snow had acquired an undiagnosed illness associated with severe fatigue that affected his work for the rest of his life. The Snows lived in Headington from 1930 to 1960, when Robin Snow resigned his fellowship for health reasons. After five years living in
Budleigh Salterton Budleigh Salterton is a seaside town on the coast in East Devon, England, south-east of Exeter. It lies within the East Devon Area of Outstanding Natural Beauty, and forms much of the electoral ward of Budleigh, whose ward population at t ...
, the Snows moved to
Vernet-les-Bains Vernet-les-Bains (; ca, Vernet) is a commune in the Pyrénées-Orientales department in southern France. It is a centre for visitors and holidaymakers. The village has a sunny climate (with, on average, 300 days of sunshine each year) and is s ...
in the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to ...
. Robin Snow died on 1 August 1969. Mary Snow died in 1978 in Perpignan. They had no children. Snow and her husband both enjoyed rock climbing and investigation of the
paranormal Paranormal events are purported phenomena described in popular culture, folk, and other non-scientific bodies of knowledge, whose existence within these contexts is described as being beyond the scope of normal scientific understanding. Not ...
. Mary Snow's work was largely carried out in collaboration with Robin Snow. As his research student, she worked on the regeneration of stem-apices after splitting. This and their work tested the hypotheses of Hofmeister and van Iterson that new primordia would arise in the largest gaps left by previous ones, and largely relied on dissection of the growing stem of ''
Lupinus albus ''Lupinus albus'', commonly known as the white lupin or field lupine, is a member of the genus '' Lupinus'' in the family Fabaceae. It is a traditional pulse cultivated in the Mediterranean region. Description The white lupin is annual, mo ...
'' under a microscope. In all the work published by them jointly, Mary Snow carried out almost all of the practical manipulation, they shared the interpretation of the results and the generation of new experimental ideas, while Robin alone wrote them up. All of the experimental work was carried out in their home, Southerway, in
Headington Headington is an eastern suburb of Oxford, England. It is at the top of Headington Hill overlooking the city in the Thames valley below, and bordering Marston to the north-west, Cowley to the south, and Barton and Risinghurst to the east. Th ...
, whether the couple frequently entertained undergraduates and research students. The Snows published a series of papers on the subject from 1931 onwards, which are still widely cited. Mary Snow's benefaction to the
Oxford Botanic Gardens The University of Oxford Botanic Garden is the oldest botanic garden in Great Britain and one of the oldest scientific gardens in the world. The garden was founded in 1621 as a physic garden growing plants for medicinal research. Today it conta ...
made possible the development of their arboretum at
Nuneham Courtenay Nuneham Courtenay is a village and civil parish about southeast of Oxford. It occupies a pronounced section of the left bank of the River Thames. Geography The parish is bounded to the west by the River Thames and on other sides by field bound ...
.


Selected publications

* Snow, M., & Snow, GRS. (1931). Experiments on Phyllotaxis. I. The Effect of Isolating a Primordium. ''Philosophical Transactions of the Royal Society of London. Series B,'' ''221'', 1–43. * Snow, M., & Snow, GRS. (1933). Experiments on Phyllotaxis. II. The Effect of Displacing a Primordium. ''Philosophical Transactions of the Royal Society of London. Series B'', ''222'', 353–400. * Snow, M., & Snow, GRS. (1934). The Interpretation of Phyllotaxis. ''Biological Reviews'', ''9''(1), 132–137. * Snow, M., & Snow, GRS. (1935). Experiments on Phyllotaxis. Part III. Diagonal Splits through Decussate Apices. ''Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences'', ''225''(519), 63–94. * Snow, M., & Snow, GRS. (1947). ''On the Determination of Leaves''. * Snow, M., & Snow, GRS. (1959). Regulation of sizes of leaf primordia by older leaves. ''Proceedings of the Royal Society of London. Series B. Biological Sciences'', ''151''(942), 39–47. * Snow, M., & Snow, GRS. (1962). A theory of the regulation of phyllotaxis based on Lupinus albus. ''Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences'', ''244''(717), 483–513.


References

{{DEFAULTSORT:Snow, Mary 1902 births 1978 deaths 20th-century British botanists 20th-century British women scientists People from Rainhill Alumni of St Hugh's College, Oxford Fellows of Somerville College, Oxford