A mammoth is any

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of the

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

elephantid

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

''Mammuthus'', one of the many genera that make up the

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...

of trunked

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s called

proboscidea The Proboscidea (; , ) are a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. From ...

ns. The various species of mammoth were commonly equipped with long, curved

tusks Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with pigs and walruses, or, in the case of elephants, elongated incisors. Tusks share co ...

and, in northern species, a covering of long

hair Hair is a protein filament that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals. The human body, apart from areas of glabrous skin, is covered in follicles which produce thick terminal and f ...

. They lived from the

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...

(from around 5 million years ago) into the

Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...

at about 4,000 years ago, and various species existed in Africa, Europe, Asia, and North America. They were members of the family

Elephantidae Elephantidae is a family (biology), family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are terrestrial animal, terrestrial large mammals with a snout modified into a Elephant#Trunk, trunk and teeth ...

, which also contains the two genera of modern

elephant Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae an ...

s and their ancestors. Mammoths are more closely related to living

Asian elephant The Asian elephant (''Elephas maximus''), also known as the Asiatic elephant, is the only living species of the genus ''Elephas'' and is distributed throughout the Indian subcontinent and Southeast Asia, from India in the west, Nepal in the no ...

s than African elephants. The oldest representative of ''Mammuthus'', the South African mammoth (''M. subplanifrons''), appeared around 5 million years ago during the early

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Columbian mammoth The Columbian mammoth (''Mammuthus columbi'') is an extinct species of mammoth that inhabited the Americas as far north as the Northern United States and as far south as Costa Rica during the Pleistocene epoch. It was one of the last in a line ...

(''M. columbi''). The last species to emerge, the

woolly mammoth The woolly mammoth (''Mammuthus primigenius'') is an extinct species of mammoth that lived during the Pleistocene until its extinction in the Holocene epoch. It was one of the last in a line of mammoth species, beginning with '' Mammuthus subp ...

(''M. primigenius''), developed about 400,000 years ago in East Asia, with some surviving on Russia's

Wrangel Island Wrangel Island ( rus, О́стров Вра́нгеля, r=Ostrov Vrangelya, p=ˈostrəf ˈvrangʲɪlʲə; ckt, Умӄиԓир, translit=Umqiḷir) is an island of the Chukotka Autonomous Okrug, Russia. It is the 91st largest island in the w ...

in the

Arctic Ocean The Arctic Ocean is the smallest and shallowest of the world's five major oceans. It spans an area of approximately and is known as the coldest of all the oceans. The International Hydrographic Organization (IHO) recognizes it as an ocean, a ...

, as well as possibly the

Taymyr Peninsula The Taymyr Peninsula (russian: Таймырский полуостров, Taymyrsky poluostrov) is a peninsula in the Far North of Russia, in the Siberian Federal District, that forms the northernmost part of the mainland of Eurasia. Administrati ...

on mainland

Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...

, until as recently as roughly 3,700 to 4,000 years ago, still extant during the construction of the

Great Pyramid The Great Pyramid of Giza is the biggest Egyptian pyramid and the tomb of Fourth Dynasty pharaoh Khufu. Built in the early 26th century BC during a period of around 27 years, the pyramid is the oldest of the Seven Wonders of the Ancient World, ...

of ancient Egypt.

Evolution

The earliest known

proboscideans The Proboscidea (; , ) are a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. From ...

, the

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

that contains the elephants, existed about 55 million years ago around the

Tethys Sea The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...

area. The closest relatives of the Proboscidea are the

sirenians The Sirenia (), commonly referred to as sea-cows or sirenians, are an order (biology), order of fully aquatic, herbivorous mammals that inhabit swamps, rivers, estuaries, marine wetlands, and coastal marine waters. The Sirenia currently compris ...

and the

hyrax Hyraxes (), also called dassies, are small, thickset, herbivorous mammals in the order Hyracoidea. Hyraxes are well-furred, rotund animals with short tails. Typically, they measure between long and weigh between . They are superficially simil ...

es. The family

is known to have existed six million years ago in Africa, and includes the living elephants and the mammoths. Among many now extinct clades, the

mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of th ...

is only a distant relative of the mammoths, and part of the separate

Mammutidae Mammutidae is an extinct family of proboscideans that appeared during the Oligocene epoch and survived until the start of the Holocene. The family was first described in 1922, classifying fossil specimens of the type genus ''Mammut'' (mastodons) ...

family, which diverged 25 million years before the mammoths evolved. The following

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

shows the placement of the genus ''Mammuthus'' among other proboscideans, based on

hyoid The hyoid bone (lingual bone or tongue-bone) () is a horseshoe-shaped bone situated in the anterior midline of the neck between the chin and the thyroid cartilage. At rest, it lies between the base of the mandible and the third cervical vertebra. ...

characteristics: MammothVsMastodon

Since many remains of each species of mammoth are known from several localities, it is possible to reconstruct the evolutionary history of the genus through morphological studies. Mammoth species can be identified from the number of enamel ridges on their molars; the primitive species had few ridges, and the amount increased gradually as new species evolved and replaced the former ones. At the same time, the crowns of the teeth became longer, and the skulls become higher from top to bottom and shorter from the back to the front over time to accommodate this. Mammuthus africanavus plio tunisia

The first known members of the genus ''Mammuthus'' are the African species ''

Mammuthus subplanifrons ''Mammuthus subplanifrons'', the South African Mammoth, is the oldest representative of the genus ''Mammuthus'', appearing around 5 million years ago during the early Pliocene in what is today South Africa and countries of East Africa East ...

'' from the

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Mammuthus africanavus'' from the

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...

. The former is thought to be the ancestor of later forms. Mammoths entered Europe around 3 million years ago; the earliest known type has been named ''M. rumanus'', which spread across Europe and China. Only its molars are known, which show it had 8–10 enamel ridges. A population evolved 12–14 ridges and split off from and replaced the earlier type, becoming ''

M. meridionalis ''Mammuthus meridionalis'', or the southern mammoth, is an extinct species of mammoth native to Europe and Central Asia from the Gelasian stage of the Early Pleistocene, living from 2.5–0.8 mya. Taxonomy The taxonomy of extinct elephant ...

''. In turn, this species was replaced by the steppe mammoth, ''

M. trogontherii The steppe mammoth (''Mammuthus trogontherii'', sometimes ''Mammuthus armeniacus'') is an extinct species of Elephantidae that ranged over most of northern Eurasia during the late Early and Middle Pleistocene, approximately 1.8 million-200,000 y ...

'', with 18–20 ridges, which evolved in East Asia ca. 1.8 million years ago. Steppe mammoth populations replaced ''

'' in Europe between 1 and 0.7 million years ago. The Columbian mammoth, ''

M. columbi The Columbian mammoth (''Mammuthus columbi'') is an extinct species of mammoth that inhabited the Americas as far north as the Northern United States and as far south as Costa Rica during the Pleistocene epoch. It was one of the last in a line ...

'', evolved from a population of ''M. trogontherii'' that had entered North America over 1 million years ago. Mammoths derived from ''M. trogontherii'' evolved molars with 26 ridges between 800,000 and 400,000 years ago in Siberia, becoming the

, ''M. primigenius''. The woolly mammoth would replace the steppe mammoth in Europe during the late Middle Pleistocene around 200,000 years ago. A 2011 genetic study showed that two examined specimens of the Columbian mammoth were grouped within a subclade of woolly mammoths. This suggests that the two populations interbred and produced fertile offspring. It also suggested that a North American form known as "''M. jeffersonii''" may be a hybrid between the two species. By the late Pleistocene, mammoths in continental Eurasia had undergone a major transformation, including a shortening and heightening of the cranium and mandible, increase in molar hypsodonty index, increase in plate number, and thinning of dental enamel. Due to this change in physical appearance, it became customary to group European mammoths separately into distinguishable clusters: # Early Pleistocene – ''Mammuthus meridionalis'' # Middle Pleistocene – ''Mammuthus trogontherii'' # Late Pleistocene – ''Mammuthus primigenius'' There is speculation as to what caused this variation within the three chronospecies. Variations in environment, climate change, and migration surely played roles in the evolutionary process of the mammoths. Take ''M. primigenius'' for example: Woolly mammoths lived in opened grassland biomes. The cool steppe-tundra of the Northern Hemisphere was the ideal place for mammoths to thrive because of the resources it supplied. With occasional warmings during the ice age, climate would change the landscape, and resources available to the mammoths altered accordingly. In February 2021, scientists reported that DNA from million-year old mammoth

remains Remains or The Remains may refer to: Music *The Remains (band), a 1960s American rock band *The Ramainz, originally The Remains, a Ramones tribute band Albums * ''Remains'' (Alkaline Trio album), 2007 * ''Remains'' (Annihilator album), 1997 * ' ...

had become the oldest ever

sequenced In genetics and biochemistry, sequencing means to determine the primary structure (sometimes incorrectly called the primary sequence) of an unbranched biopolymer. Sequencing results in a symbolic linear depiction known as a sequence which suc ...

Etymology and early observations

The word ''mammoth'' was first used in Europe during the early 17th century, when referring to ''maimanto'' tusks discovered in Siberia. John Bell, who was on the

Ob River } The Ob ( rus, Обь, p=opʲ: Ob') is a major river in Russia. It is in western Siberia; and together with Irtysh forms the world's List of rivers by length, seventh-longest river system, at . It forms at the confluence of the Biya (river), Biya ...

in 1722, said that mammoth tusks were well known in the area. They were called "mammon's horn" and were often found in washed-out river banks. Some local people claimed to have seen a living mammoth, but they came out only at night and always disappeared under water when detected. He bought one and presented it to Hans Sloan who pronounced it an elephant's tooth. The folklore of some native peoples of Siberia, who would routinely find mammoth bones, and sometimes frozen mammoth bodies, in eroding river banks, had various interesting explanations for these finds. Among the

Khanty people The Khanty (Khanty: ханти, ''hanti''), also known in older literature as Ostyaks (russian: остяки) are a Ugric indigenous people, living in Khanty–Mansi Autonomous Okrug, a region historically known as "Yugra" in Russia, together ...

of the

Irtysh River The Irtysh ( otk, 𐰼𐱅𐰾:𐰇𐰏𐰕𐰏, Ertis ügüzüg, mn, Эрчис мөрөн, ''Erchis mörön'', "erchleh", "twirl"; russian: Иртыш; kk, Ертіс, Ertis, ; Chinese: 额尔齐斯河, pinyin: ''É'ěrqísī hé'', Xiao'erj ...

basin, a belief existed that the mammoth was some kind of a water spirit. According to other Khanty, the mammoth was a creature that lived underground, burrowing its tunnels as it went, and would die if it accidentally came to the surface. The concept of the mammoth as an underground creature was known to the Chinese, who received some mammoth ivory from the Siberian natives; accordingly, the creature was known in China as ''yǐn shǔ'' 隐鼠, "the hidden rodent".

Thomas Jefferson Thomas Jefferson (April 13, 1743 – July 4, 1826) was an American statesman, diplomat, lawyer, architect, philosopher, and Founding Fathers of the United States, Founding Father who served as the third president of the United States from 18 ...

, who famously had a keen interest in

paleontology Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

, is partially responsible for transforming the word ''mammoth'' from a noun describing the prehistoric elephant to an adjective describing anything of surprisingly large size. The first recorded use of the word as an adjective was in a description of a large wheel of cheese (the "

Cheshire Mammoth Cheese The Cheshire Mammoth Cheese was a gift from the town of Cheshire, Massachusetts to President of the United States of America, President Thomas Jefferson in 1802. The cheese was created by combining the milk from every cow in the town, and made in ...

") given to Jefferson in 1802.Simpson, J. (2009).
Word Stories: Mammoth
" ''Oxford English Dictionary Online'', Oxford University Press. Accessed 05-JUN-2009.

Description

Like their modern relatives, mammoths were quite large. The largest known species reached heights in the region of at the shoulder and weights of up to , while exceptionally large males may have exceeded . However, most species of mammoth were only about as large as a modern

(which are about 2.5 m to 3 m high at the shoulder, and rarely exceeding 5 tonnes). Both sexes bore tusks. A first, small set appeared at about the age of six months, and these were replaced at about 18 months by the permanent set. Growth of the permanent set was at a rate of about per year. Mammuthus Size comparison

Based on studies of their close relatives, the modern elephants, mammoths probably had a

gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregna ...

period of 22 months, resulting in a single calf being born. Their social structure was probably the same as that of

African African or Africans may refer to: * Anything from or pertaining to the continent of Africa: ** People who are native to Africa, descendants of natives of Africa, or individuals who trace their ancestry to indigenous inhabitants of Africa *** Ethn ...

and Asian elephants, with females living in herds headed by a matriarch, whilst bulls lived solitary lives or formed loose groups after sexual maturity. Scientists discovered and studied the remains of a mammoth calf, and found that fat greatly influenced its form, and enabled it to store large amounts of nutrients necessary for survival in temperatures as low as . The fat also allowed the mammoths to increase their muscle mass, allowing the mammoths to fight against enemies and live longer. Woolly mammoths evolved a suite of adaptations for arctic life, including morphological traits such as small ears and tails to minimize heat loss, a thick layer of subcutaneous fat, long thick fur, and numerous sebaceous glands for insulation , as well as a large brown-fat hump like deposit behind the neck that may have functioned as a heat source and fat reservoir during winter.

Diet

Depending on the species or race of mammoth, the diet differed somewhat depending on location, although all mammoths ate similar things. For the Columbian mammoth, ''

'', the diet was mainly

grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and other ...

. American Columbian mammoths fed primarily on cactus leaves, trees, and shrubs. These assumptions were based on mammoth feces and mammoth teeth. Mammoths, like modern day elephants, have hypsodont molars. These features also allowed mammoths to live an expansive life because of the availability of grasses and trees. For the Mongochen mammoth, its diet consisted of herbs, grasses, larch, and shrubs, and possibly

alder Alders are trees comprising the genus ''Alnus'' in the birch family Betulaceae. The genus comprises about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few sp ...

. These inferences were made through the observation of mammoth feces, which scientists observed contained non-arboreal pollen and moss spores. European mammoths had a major diet of

C3 carbon fixation carbon fixation is the most common of three metabolic pathways for carbon fixation in photosynthesis, along with and CAM. This process converts carbon dioxide and ribulose bisphosphate (RuBP, a 5-carbon sugar) into two molecules of 3-phosph ...

plants. This was determined by examining the isotopic data from the European mammoth teeth. The arctic tundra and steppe where the mammoths lived appears to have been dominated by

forb A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...

s, not grass. These were richer in protein and easier to digest than grasses and wooden plants, which came to dominate the areas when the climate became wetter and warmer. This could have been a major contributor to why the arctic megafauna went extinct. The Yamal baby mammoth Lyuba, found in 2007 in the Yamal Peninsula in Western Siberia, suggests that baby mammoths, as do modern baby elephants, ate the dung of adult animals. The evidence to show this is that the dentition (teeth) of the baby mammoth had not yet fully developed to chew grass. Furthermore, there was an abundance of ascospores of

coprophilous fungi Coprophilous fungi (''dung-loving'' fungi) are a type of saprobic fungi that grow on animal dung. The hardy spores of coprophilous species are unwittingly consumed by herbivores from vegetation, and are excreted along with the plant matter. The f ...

from the pollen spectrum of the baby's mother. Coprophilous fungi are fungi that grow on animal dung and disperse spores in nearby vegetation, which the baby mammoth would then consume. Spores might have gotten into its stomach while grazing for the first few times. Coprophagy may be an

adaptation In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...

, serving to populate the infant's gut with the needed

microbiome A microbiome () is the community of microorganisms that can usually be found living together in any given habitat. It was defined more precisely in 1988 by Whipps ''et al.'' as "a characteristic microbial community occupying a reasonably well ...

for digestion. Mammoths alive in the

Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...

during the

Last Glacial Maximum The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent. Ice sheets covered much of Northern North America, Northern Eur ...

consumed mainly

s, such as ''

Artemisia Artemisia may refer to: People * Artemisia I of Caria (fl. 480 BC), queen of Halicarnassus under the First Persian Empire, naval commander during the second Persian invasion of Greece * Artemisia II of Caria (died 350 BC), queen of Caria under th ...

'';

graminoid In botany and ecology, graminoid refers to a herbaceous plant with a grass-like morphology, i.e. elongated culms with long, blade-like leaves. They are contrasted to forbs, herbaceous plants without grass-like features. The plants most ofte ...

s were only a minor part of their diet.

Extinction

The

(''M. primigenius'') was the last species of the genus. Most populations of the woolly mammoth in North America and Eurasia, as well as all the

Columbian mammoth The Columbian mammoth (''Mammuthus columbi'') is an extinct species of mammoth that inhabited the Americas as far north as the Northern United States and as far south as Costa Rica during the Pleistocene epoch. It was one of the last in a line ...

s (''M. columbi'') in North America, died out around the time of the last

glacial retreat The retreat of glaciers since 1850 affects the availability of fresh water for irrigation and domestic use, mountain recreation, animals and plants that depend on glacier-melt, and, in the longer term, the level of the oceans. Deglaciation occu ...

, as part of a

mass extinction An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. It ...

megafauna In terrestrial zoology, the megafauna (from Greek μέγας ''megas'' "large" and New Latin ''fauna'' "animal life") comprises the large or giant animals of an area, habitat, or geological period, extinct and/or extant. The most common threshold ...

in northern Eurasia and the Americas. Until recently, the last woolly mammoths were generally assumed to have vanished from Europe and southern Siberia about 12,000 years ago, but new findings show some were still present there about 10,000 years ago. Slightly later, the woolly mammoths also disappeared from continental northern Siberia. A small population survived on

St. Paul Island, Alaska Saint Paul Island (russian: Остров Святого Павла, Ostrov Svyatogo Pavla) is the largest of the Pribilof Islands, a group of four Alaskan volcanic islands located in the Bering Sea between the United States and Russia. The city o ...

, up until 3750 BC, and the small mammoths of

survived until about 2000 BC Recent eDNA research of sediments indicates mammoths survived in north central Siberia at least as late as 2000 BC, in continental northeast Siberia until at least 5300 BC, and until at least 6600 BC in North America. The warming trend (Holocene) that occurred 12,000 years ago, accompanied by a glacial retreat and rising sea levels, has been suggested as a contributing factor. Forests replaced open woodlands and grasslands across the continent. The available habitat would have been reduced for some megafaunal species, such as the mammoth. However, such climate changes were nothing new; numerous very similar warming episodes had occurred previously within the

ice age An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages and gree ...

of the last several million years without producing comparable megafaunal extinctions, so climate alone is unlikely to have played a decisive role. The spread of advanced human hunters through northern

Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...

and the

Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...

around the time of the extinctions, however, was a new development, and thus might have contributed significantly. Hebior Mammoth Clean

Whether the general mammoth population died out for climatic reasons or due to

overhunting Overexploitation, also called overharvesting, refers to harvesting a renewable resource to the point of diminishing returns. Continued overexploitation can lead to the destruction of the resource, as it will be unable to replenish. The term app ...

by humans is controversial. During the transition from the Late Pleistocene epoch to the Holocene epoch, there was shrinkage of the distribution of the mammoth because progressive warming at the end of the Pleistocene epoch changed the mammoth's environment. The

mammoth steppe During the Last Glacial Maximum, the mammoth steppe, also known as steppe-tundra, was the Earth's most extensive biome. It spanned from Spain eastward across Eurasia to Canada and from the List of islands in the Arctic Ocean, arctic islands sout ...

was a periglacial landscape with rich herb and grass vegetation that disappeared along with the mammoth because of environmental changes in the climate. Mammoths had moved to isolated spots in Eurasia, where they disappeared completely. Also, it is thought that Late Paleolithic and Mesolithic human hunters might have affected the size of the last mammoth populations in Europe. There is evidence to suggest that humans did cause the mammoth extinction, although there is no definitive proof. It was found that humans living south of a mammoth steppe learned to adapt themselves to the harsher climates north of the steppe, where mammoths resided. It was concluded that if humans could survive the harsh north climate of that particular mammoth steppe then it was possible humans could hunt (and eventually extinguish) mammoths everywhere. Another hypothesis suggests mammoths fell victim to an infectious disease. A combination of

climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...

and hunting by humans may be a possible explanation for their extinction. ''

Homo erectus ''Homo erectus'' (; meaning "upright man") is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Several human species, such as '' H. heidelbergensis'' and '' H. antecessor' ...

'' is known to have consumed mammoth meat as early as 1.8 million years ago, though this may mean only successful scavenging, rather than actual hunting. Later humans show greater evidence for hunting mammoths; mammoth bones at a 50,000-year-old site in South Britain suggest that

Neanderthals Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an Extinction, extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ag ...

butchered the animals, while various sites in Eastern Europe dating from 15,000 to 44,000 years old suggest humans (probably ''

Homo sapiens Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...

'') built dwellings using mammoth bones (the age of some of the earlier structures suggests that Neanderthals began the practice). However, the

American Institute of Biological Sciences The American Institute of Biological Sciences (AIBS) is a nonprofit scientific charity. The organization’s mission is to promote the use of science to inform decision-making and advance biology for the benefit of science and society. Overvie ...

notes that bones of dead elephants, left on the ground and subsequently trampled by other elephants, tend to bear marks resembling butchery marks, which have allegedly been misinterpreted as such by

archaeologists Archaeology or archeology is the scientific study of human activity through the recovery and analysis of material culture. The archaeological record consists of artifacts, architecture, biofacts or ecofacts, sites, and cultural landscap ...

. In 2021, a study using ancient

environmental DNA Environmental DNA or eDNA is DNA that is collected from a variety of environmental samples such as soil, seawater, snow or air, rather than directly sampled from an individual organism. As various organisms interact with the environment, DNA ...

concluded that the extinction of the mammoth was primarily caused by dramatic vegetation changes at the end of the

, due to a changed climate and precipitation regime. During the Last Glacial Maximum, the Arctic would have had a homogeneous flora consisting of mammoth steppe-associated plants, supporting a unique community of grazing mammals, including the mammoth. Increased precipitation following the end of the LGM caused caused a significant

paludification Paludification is the most common process by which peatlands in the boreal zone are formed. Formation The process is characterized by peat initialization on previously drier and vegetated habitats over inorganic soils, with no fully aquatic ...

of the steppe, which has been linked to a general decrease in species diversity of the region's former inhabitants, and not just mammoths. In addition, the study found mammoths to have persisted on the mainland for far longer than previously thought, with their shrinking distribution roughly tracking the shrinkage of the mammoth steppe; their final mainland refugium, the

, was roughly concurrent with the Wrangel population. The close relationship between decreasing mammoth populations and steppe, as well as the lack of a sudden decline that would be associated with the human overkill hypothesis, indicates that humans and mammoths may have coexisted for millennia, and thus humans may have only played a minor role in the species' extinction. Many hypotheses also seek to explain the regional extinction of mammoths in specific areas. Scientists have speculated that the mammoths of Saint Paul Island (Alaska), an isolated enclave where mammoths survived until about 8,000 years ago, died out as the island shrank by 80–90% when sea levels rose, eventually making it too small to support a viable population. Similarly, genome sequences of the Wrangel Island mammoths indicate a sharp decline in genetic diversity, though the extent to which this played a role in their extinction is still unclear. Another hypothesis, said to be the cause of mammoth extinction in Siberia, comes from the idea that many may have drowned. While traveling to the Northern River, many of these mammoths broke through the ice and drowned. This also explains bones remains in the Arctic Coast and some of the

New Siberian Islands The New Siberian Islands ( rus, Новосиби́рские Oстрова, r=Novosibirskiye Ostrova; sah, Саҥа Сибиир Aрыылара, translit=Saña Sibiir Arıılara) are an archipelago in the Extreme North of Russia, to the north o ...

Dwarfing Dwarfing is a process in which a breed of animals or cultivar of plants is changed to become significantly smaller than standard members of their species. The effect can be induced through human intervention or non-human processes, and can include ...

occurred with the

pygmy mammoth The pygmy mammoth or Channel Islands mammoth (''Mammuthus exilis'') is an extinct species of dwarf elephant descended from the Columbian mammoth (''M. columbi'') of mainland North America. This species became extinct during the Quaternary extin ...

on the outer

Channel Islands of California The Channel Islands () are an eight-island archipelago located within the Southern California Bight in the Pacific Ocean, off the coast of California. The four Northern Channel Islands are part of the Transverse Ranges geologic province, and ...

, but at an earlier period. Those animals were very likely killed by early Paleo-Native Americans, and habitat loss caused by a rising sea level that split

Santa Rosae Santa Rosae (also spelled Santarosae) was, before the end of the last ice age, an ancient landmass off the coast of present-day southern California, near Santa Barbara County and Ventura County, of which the northern Channel Islands of California ...

into the outer Channel Islands.

Mammoth-elephant hybrid

One proposed scientific use of this preserved genetic material is to recreate living mammoths. This has long been discussed theoretically but has only recently become the subject of formal effort due to advances in

molecular biology Molecular biology is the branch of biology that seeks to understand the molecular basis of biological activity in and between cells, including biomolecular synthesis, modification, mechanisms, and interactions. The study of chemical and physi ...

techniques and cloning of mammals.The Long Now Fouldation - Revive and Restore
According to one research team, a mammoth cannot be recreated, but they will try to eventually grow in an "artificial womb" a hybrid elephant with some woolly mammoth traits.The Mammoth Genome Project

Pennsylvania State University The Pennsylvania State University (Penn State or PSU) is a Public university, public Commonwealth System of Higher Education, state-related Land-grant university, land-grant research university with campuses and facilities throughout Pennsylvan ...

. Accessed: October 2018. The outcome would be an elephant-mammoth hybrid with no more than 1% mammoth genes. Other projects are working on gradually adding mammoth genes to

cells ''in vitro''."Mammoth: Back from the Dead"
. ''National Geographic Channel''.

Evolution

Etymology and early observations

Description

Diet

Extinction

Mammoth-elephant hybrid

See also

References

Further reading