Malus Florina 4485
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''Malus'' ( or ) is a genus of about 30–55 species of small deciduous trees or
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s in the family
Rosaceae Rosaceae (), the rose family, is a medium-sized family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus ''Rosa''. Among the most species-rich genera are ''Alchemilla'' (270), ''Sorbus ...
, including the domesticated orchard apple, crab apples, wild apples, and rainberries. The genus is native to the temperate zone of the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
.


Description

Apple trees are typically talI at maturity, with a dense, twiggy crown. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are long, alternate, simple, with a serrated margin. The flowers are borne in corymbs, and have five
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s, which may be white, pink, or red, and are
perfect Perfect commonly refers to: * Perfection, completeness, excellence * Perfect (grammar), a grammatical category in some languages Perfect may also refer to: Film * Perfect (1985 film), ''Perfect'' (1985 film), a romantic drama * Perfect (2018 f ...
, with usually red stamens that produce copious
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
, and a half-inferior ovary; flowering occurs in the spring after 50–80 growing degree days (varying greatly according to subspecies and cultivar). Many apples require cross-pollination between individuals by insects (typically
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyly, monophyletic lineage within the ...
s, which freely visit the flowers for both
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
and
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
); these are called self-sterile, so self-pollination is impossible, making pollinating insects essential. A number of cultivars are self-pollinating, such as 'Granny Smith' and 'Golden Delicious', but are considerably fewer in number compared to their cross-pollination dependent counterparts. Several ''Malus'' species, including domestic apples, hybridize freely. The fruit is a globose
pome In botany, a pome is a type of fruit produced by flowering plants in the subtribe Malinae of the family Rosaceae. Well-known pomes include the apple, pear, and quince. Etymology The word ''pome'' entered English in the late 14th century, and re ...
, varying in size from in diameter in most of the wild species, to in ''M. sylvestris sieversii'', in ''M. domestica'', and even larger in certain cultivated orchard apples. The centre of the fruit contains five carpels arranged star-like, each containing one or two seeds.


Subdivisions and species

About 42 to 55 species and natural hybrids are known, with about 25 from China, of which 15 are endemic. The genus ''Malus'' is subdivided into eight sections (six, with two added in 2006 and 2008). The genus '' Docynia'' has been shown to be nested within ''Malus'' in molecular phylogenies. The oldest fossils of the genus date to the Eocene ( Lutetian), which are leaves belonging to the species ''Malus collardii'' and ''Malus kingiensis'' from western North America (Idaho) and the Russian Far East ( Kamchatka), respectively.


Natural hybrids

* ''
Malus × micromalus ''Malus'' × ''micromalus'', the midget crabapple or Kaido crabapple, is a species in the genus ''Malus ''Malus'' ( or ) is a genus of about 30–55 species of small deciduous trees or shrubs in the family Rosaceae, including the domestica ...
'' – midget crabapple


Fossil species

After * ''Malus collardii'' Axelrod, North America (Idaho), Eocene * ''Malus kingiensis'' Budants, Kamchatka Peninsula, Russia, Eocene * ''Malus florissantensis'' (Cockerell) MacGinitie Green River Formation, North America (Colorado) Eocene * ''Malus pseudocredneria'' (Cockerell) MacGinitie Green River Formation, North America (Colorado) Eocene * ''Malus idahoensis'' R.W.Br. North America (Idaho), Miocene * ''Malus parahupehensis'' J.Hsu and R.W.Chaney Shanwang, Shandong, China, Miocene * ''Malus antiqua'' Doweld Romania, Pliocene * ''Malus pseudoangustifolia'' E.W.Berry North America (South Carolina), Pleistocene


Cultivation

Crabapples are popular as compact ornamental trees, providing blossom in spring and colourful fruit in autumn. The fruits often persist throughout winter. Numerous hybrid cultivars have been selected. Some crabapples are used as rootstocks for domestic apples to add beneficial characteristics. For example, the rootstocks of '' Malus baccata'' varieties are used to give additional cold hardiness to the combined plants for orchards in cold northern areas. They are also used as pollinizers in apple orchards. Varieties of crabapple are selected to bloom contemporaneously with the apple variety in an orchard planting, and the crabs are planted every sixth or seventh tree, or limbs of a crab tree are grafted onto some of the apple trees. In emergencies, a bucket or drum bouquet of crabapple flowering branches is placed near the beehives as orchard pollenizers. Because of the plentiful blossoms and small fruit, crabapples are popular for use in
bonsai Bonsai ( ja, 盆栽, , tray planting, ) is the Japanese art of growing and training miniature trees in pots, developed from the traditional Chinese art form of ''penjing''. Unlike ''penjing'', which utilizes traditional techniques to produce ...
culture.


Cultivars

These cultivars have won the Royal Horticultural Society's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
:- *'Adirondack' *'Butterball' * 'Comtesse de Paris' *'Evereste' *'Jelly King'='Mattfru' *'Laura' *'' Malus × robusta'' 'Red Sentinel' *'Sun Rival' Other varieties are dealt with under their species names.


Toxicity

The seeds contain
cyanide Cyanide is a naturally occurring, rapidly acting, toxic chemical that can exist in many different forms. In chemistry, a cyanide () is a chemical compound that contains a functional group. This group, known as the cyano group, consists of a ...
compounds.


Uses

Crabapple fruit is not an important crop in most areas, being extremely sour due to malic acid (which like the genus derives from the Latin name '' mālum''), and in some species woody, so is rarely eaten raw. In some Southeast Asian cultures, they are valued as a sour condiment, sometimes eaten with salt and chilli or shrimp paste. Some crabapple varieties are an exception to the reputation of being sour, and can be very sweet, such as the 'Chestnut' cultivar. Crabapples are an excellent source of
pectin Pectin ( grc, πηκτικός ': "congealed" and "curdled") is a heteropolysaccharide, a structural acid contained in the primary lamella, in the middle lamella, and in the cell walls of terrestrial plants. The principal, chemical component of ...
, and their juice can be made into a ruby-coloured preserve with a full, spicy flavour. A small percentage of crabapples in
cider Cider ( ) is an alcoholic beverage made from the fermented juice of apples. Cider is widely available in the United Kingdom (particularly in the West Country) and the Republic of Ireland. The UK has the world's highest per capita consumption, ...
makes a more interesting flavour. As
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlement of Britain, Anglo ...
''Wergulu'', the crab apple is one of the nine plants invoked in the pagan Anglo-Saxon '' Nine Herbs Charm'', recorded in the 10th century. Applewood gives off a pleasant scent when burned, and smoke from an applewood fire gives an excellent flavour to
smoked Smoking is the process of flavoring, browning, cooking, or preserving food by exposing it to smoke from burning or smoldering material, most often wood. Meat, fish, and ''lapsang souchong'' tea are often smoked. In Europe, alder is the tradi ...
foods. It is easier to cut when green; dry applewood is exceedingly difficult to carve by hand. It is a good wood for cooking fires because it burns hot and slow, without producing much flame.


Gallery

File:Apple blossom. Eastern Siberia.jpg, Apple blossom. Eastern Siberia File:Crabapples.jpg, Ripe wild crab apples (''Malus sylvestris'') File:Lee-Russell-Farm-Security-Administration-1939-Crab-Apples.jpg, Baskets of crab apples for sale in Connecticut in 1939 File:20071103Tradkrafta1.JPG, Trunk


References


External links


Germplasm Resources Information Network: ''Malus''

Flora of China: ''Malus''




a cooperative among Purdue University, Rutgers University, and the University of Illinois. {{Authority control Rosaceae genera Plants used in bonsai Fruit trees Taxa named by Philip Miller