Malthininae
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The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are cosmopolitan in distribution. One of the first described species has a color pattern reminiscent of the red coats of early British soldiers, hence the common name. They are also known commonly as leatherwings because of their soft elytra. Historically, these beetles were placed in a superfamily "Cantharoidea", which has been subsumed by the superfamily Elateroidea; the name is still sometimes used as a rankless grouping, including the families Cantharidae, Lampyridae,
Lycidae The Lycidae are a family in the beetle order Coleoptera, members of which are commonly called net-winged beetles. These beetles are cosmopolitan, being found in Nearctic, Palearctic, Neotropical, Afrotropical, Oriental, and Australian ecoregions ...
, Omethidae (which includes
Telegeusidae Omethidae is a family of Elateroidea sometimes known as the false soldier beetles. They are native to South, Southeast and Eastern Asia and the Americas. Their biology is obscure and their larvae are unknown. They appear to inhabit vegetation in ...
), Phengodidae, and Rhagophthalmidae. Soldier beetles often feed on both nectar and pollen as well as predating other small insects. The larvae are often active, velvety, often brightly-colored, and they feed on the ground, hunting snails and other small creatures.


Evolutionary history

The oldest described member of the family is '' Molliberus'' from the
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
(early Albian) aged El Soplao amber from Cantabria, Spain, belonging to the tribe
Cantharini Cantharini is a tribe of soldier beetles in the family Cantharidae. There are more than 40 genera and over 500 described species in Cantharini. Genera These 46 genera belong to the tribe Cantharini: * '' Ancistronycha'' Märkel, 1852 * '' Arm ...
in the subfamily
Cantharinae Cantharinae is a subfamily of beetles in the family Cantharidae. There are at least 200 described species in Cantharinae. ;Taxonomic note: *Lawrence and Newton (1995) give the authorship of this subfamily as Imhoff, 1856 (1815). Tribes and gene ...
. Other described genera include 6 from the early Late Cretaceous (early
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
) aged Burmese amber, with 5 belonging to Cantharinae and one to Malthininae, and '' Katyacantharis'', from the Cenomanian aged Agdzhakend amber of Azerbaijan, suggested to belong to Cantharinae. Indeterminate specimens have been reported from the
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
aged Koonwarra fossil bed of the
Strzelecki Group The Halibut Field, is an oil field, within the Gippsland Basin. The oil field is located approximately 64 km offshore of southeastern Australia. The total area of this field is 26.9 km2 and is composed of 10 mappable units. Geologic h ...
, Australia and the Barremian aged Lebanese amber.


Subfamilies, tribes and selected genera

Five subfamilies are normally accepted:


Cantharinae Cantharinae is a subfamily of beetles in the family Cantharidae. There are at least 200 described species in Cantharinae. ;Taxonomic note: *Lawrence and Newton (1995) give the authorship of this subfamily as Imhoff, 1856 (1815). Tribes and gene ...

* tribe Cantharini ** '' Cantharis'' ** '' Rhagonycha'' * tribe Podabrini ** ''
Podabrus ''Podabrus'' is a genus of soldier beetles in the family Cantharidae. There are at least 110 described species in ''Podabrus'', recorded from Europe, North America and Asia. Many ''Podabrus'' appear to mimic MIMIC, known in capitalized form o ...
''


Chauliognathinae Chauliognathinae is a subfamily of soldier beetles in the family Cantharidae. There are at least 4 genera and at least 20 described species in Chauliognathinae. Genera * ''Belotus'' Gorham, 1881 * ''Chauliognathus ''Chauliognathus'' is a ge ...

* tribe Chauliognathini ** '' Belotus'' ** '' Chauliognathus'' * tribe Ichtyurini ** ''
Trypherus ''Trypherus'' is a genus of soldier beetles in the family Cantharidae The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are cosmopolitan in distribution. One of the first described species has a color ...
''


Dysmorphocerinae

* '' Afronycha'' Wittmer, 1949 - central-southern Africa * '' Asilis'' Broun, 1893 - New Zealand * '' Compsonycha'' * '' Dysmorphocerus'' Solier, 1849 * '' Flabelloontelus'' * '' Geigyella'' Wittmer, 1972 - New Guinea * '' Hansasilis'' * '' Heteromastix'' Boheman, 1858 - Australia * '' Hyponotum'' * '' Micronotum'' * '' Neoontelus'' Wittmer, 1972 - New Zealand * '' Oontelus'' Solier, 1849 - S. America * '' Plectocephalon'' * '' Plectonotum'' Gorham, 1891 -
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...


Malthininae

* tribe Malchinini ** ''
Macrocerus ''Macrocerus'' is a European genus of soldier beetles in the subfamily Malthininae and sometimes placed in the monotypic tribe Malchinini. Species ''BioLib'' lists the following species: * ''Macrocerus carinicollis'' (Weise, 1895) * ''Macroc ...
'' Motschulsky, 1845 - Europe (synonym ''Malchinus'') * tribe Malthinini ** '' Caccodes'' Sharp, 1885 - Central America, Pacific islands ** '' Malthinellus'' Kiesenwetter, 1874 - Japan ** ''
Malthinus ''Malthinus'' is a genus of soldier beetles in the family Cantharidae The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are cosmopolitan in distribution. One of the first described species has a color p ...
'' Latreille, 1805 - Japan, Europe, N. America * tribe Malthodini ** '' Frostia'' Bert. ex Guill. ** '' Malthodes'' Kiesenwetter, 1852 - mostly Europe, N. America & New Zealand ** †'' Archaeomalthodes'' Hsiao et al. 2016 Burmese amber, Myanmar, Cenomanian


Silinae

* tribe Silini ** ''
Cordylocera ''Cordylocera'' is a genus of soldier beetles (insects in the family Cantharidae). Species * ''Cordylocera albovittata'' (Wittmer, 1964) * ''Cordylocera antennata'' Guérin-Méneville, 1830 * ''Cordylocera atricornis'' Guérin-Méneville, 183 ...
'' Guérin-Méneville, 1830 ** '' Silis'' Charpentier, 1825 * tribe Tytthonyxini ** '' Tytthonyx'' LeConte, 1851


Reproduction

Large males of the soldier beetle exercise choice for larger females. Body size correlates with the abilities of males to secure females, and of females to evade males.


See also

*
List of Cantharidae genera These 115 genera belong to the family Cantharidae, soldier beetles. There are at least 1,300 described species in Cantharidae. Cantharidae genera * '' Absidia'' Mulsant, 1863 * '' Absidiella'' Wittmer, 1972 * '' Afronycha'' Wittmer, 1949 * '' An ...


References


External links


Delta key

Keys for the identification of British Cantharidae
* * {{Authority control Biological pest control beetles * Articles containing video clips