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Malesia is a biogeographical region straddling the Equator and the boundaries of the
Indomalayan The Indomalayan realm is one of the eight biogeographic realms. It extends across most of South and Southeast Asia and into the southern parts of East Asia. Also called the Oriental realm by biogeographers, Indomalaya spreads all over the Indi ...
and
Australasian realm The Australasian realm is a biogeographic realm that is coincident with, but not (by some definitions) the same as, the geographical region of Australasia. The realm includes Australia, the island of New Guinea (comprising Papua New Guinea and th ...
s, and also a phytogeographical
floristic region A phytochorion, in phytogeography, is a geographic area with a relatively uniform composition of plant species. Adjacent phytochoria do not usually have a sharp boundary, but rather a soft one, a transitional area in which many species from both r ...
in the
Paleotropical Kingdom The Paleotropical Kingdom (Paleotropis) is a floristic kingdom comprising tropical areas of Africa, Asia and Oceania (excluding Australia and New Zealand), as proposed by Ronald Good and Armen Takhtajan. Part of its flora, inherited from the ancient ...
. It has been given different definitions. The
World Geographical Scheme for Recording Plant Distributions The World Geographical Scheme for Recording Plant Distributions (WGSRPD) is a biogeographical system developed by the international Biodiversity Information Standards (TDWG) organization, formerly the International Working Group on Taxonomic Da ...
split off
Papuasia Papuasia is a Level 2 botanical region defined in the World Geographical Scheme for Recording Plant Distributions (WGSRPD). It lies in the southwestern Pacific Ocean, in the Melanesia ecoregion of Oceania and Tropical Asia. It comprises the ...
in its 2001 version.


Floristic province

Malesia was first identified as a floristic region that included the Malay Peninsula, the Malay Archipelago,
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...
, and the Bismarck Archipelago, based on a shared tropical flora derived mostly from
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
but also with numerous elements of the
Antarctic flora Antarctic flora are a distinct community of vascular plants which evolved millions of years ago on the supercontinent of Gondwana. Presently, species of Antarctica flora reside on several now separated areas of the Southern Hemisphere, includin ...
, including many species in the southern conifer families
Podocarpaceae Podocarpaceae is a large family of mainly Southern Hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Pr ...
and
Araucariaceae Araucariaceae – also known as araucarians – is an extremely ancient family of coniferous trees. The family achieved its maximum diversity during the Jurassic and Cretaceous periods and the early Cenozoic, when it was distributed almost worldw ...
. The floristic region overlaps four distinct mammalian faunal regions. The first edition of the
World Geographical Scheme for Recording Plant Distributions The World Geographical Scheme for Recording Plant Distributions (WGSRPD) is a biogeographical system developed by the international Biodiversity Information Standards (TDWG) organization, formerly the International Working Group on Taxonomic Da ...
(WGSRPD) used this definition, but in the second edition of 2001, New Guinea and the Bismarck Archipelago were removed from Malesia and were united with the
Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capit ...
, previously placed in the WGSRPD's Southwestern Pacific region, being placed into a new region,
Papuasia Papuasia is a Level 2 botanical region defined in the World Geographical Scheme for Recording Plant Distributions (WGSRPD). It lies in the southwestern Pacific Ocean, in the Melanesia ecoregion of Oceania and Tropical Asia. It comprises the ...
.


Sundaland

The western part of Malesia, which includes the Malay Peninsula and the islands of Sumatra,
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mos ...
, Bali, and
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and ea ...
, shares the large mammal fauna of Asia and is known as Sundaland. These islands are on Asia's relatively shallow continental shelf, and were linked to Asia during the
ice age An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages and gre ...
s, when sea levels were lower. The eastern edge of Sundaland is the
Wallace line The Wallace Line or Wallace's Line is a faunal boundary line drawn in 1859 by the British naturalist Alfred Russel Wallace and named by English biologist Thomas Henry Huxley that separates the biogeographical realms of Asia and Wallacea, a trans ...
, named after Alfred Russel Wallace, the nineteenth-century British naturalist who noted the difference in fauna between islands on either side of the line.
Dipterocarp Dipterocarpaceae is a family of 16 genera and about 695 known species of mainly tropical lowland rainforest trees. The family name, from the type genus ''Dipterocarpus'', is derived from Greek (''di'' = two, ''pteron'' = wing and ''karpos'' = f ...
s are predominant trees in the lowland forests of Sundaland. Sundaland has the greatest diversity of Dipterocarp species, with 10 to 14 native genera and approximately 450 native species, including approximately 267 species on Borneo, 155 on the Malay Peninsula, and 106 on Sumatra.G. Maury-Lechon and L. Curtet (1998). "Biogeography and Evolutionary Systematics of Dipterocarpaceae". in ''A Review of Dipterocarps: Taxonomy, ecology and silviculture''. Simmathiri Appanah & Jennifer M. Turnbull, eds. Center for International Forestry Research, 1998, Bogor Indonesia.


Philippines

Most of the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
were never connected to the Asian mainland, and have a largely Asian-derived flora, and a distinct mammalian fauna. The Philippines have approximately 50 species of Dipterocarps in 11 genera.


Wallacea

The islands between Sundaland and New Guinea, called
Wallacea Wallacea is a biogeographical designation for a group of mainly Indonesian islands separated by deep-water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as ...
, were never linked to the neighboring continents, and have a flora and fauna that include Indomalayan and Australasian elements. Dipterocarps, which are dominant in Sundaland, are less common in Wallacea, with only 13 species in 4 genera.


New Guinea and the Bismarck Archipelago (Sahulland)

Together with the Solomon Islands, New Guinea and the Bismarck Archipelago are placed in
Papuasia Papuasia is a Level 2 botanical region defined in the World Geographical Scheme for Recording Plant Distributions (WGSRPD). It lies in the southwestern Pacific Ocean, in the Melanesia ecoregion of Oceania and Tropical Asia. It comprises the ...
rather than Malesia in the second version of the WGSRPD. The eastern end of the earlier definition of Malesia, which includes New Guinea and the
Aru Islands The Aru Islands Regency ( id, Kabupaten Kepulauan Aru) is a group of about 95 low-lying islands in the Maluku Islands of eastern Indonesia. It also forms a regency of Maluku Province, with a land area of . At the 2011 Census the Regency had a ...
of eastern Indonesia, is linked to Australia by a shallow continental shelf, and shares many
marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a ...
mammal and bird
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
with Australia. New Guinea also has many additional elements of the Antarctic flora, including southern beech (''
Nothofagus ''Nothofagus'', also known as the southern beeches, is a genus of 43 species of trees and shrubs native to the Southern Hemisphere in southern South America (Chile, Argentina) and Australasia (east and southeast Australia, New Zealand, New Gui ...
'') and
eucalypt Eucalypt is a descriptive name for woody plants with capsule fruiting bodies belonging to seven closely related genera (of the tribe Eucalypteae) found across Australasia: ''Eucalyptus'', ''Corymbia'', ''Angophora'', '' Stockwellia'', ''Allosyn ...
s. New Guinea has the highest mountains in Malesia and Papuasia, and vegetation ranges from tropical lowland forest to tundra.


Assembly and origins of the Malesian flora

Major contributions to rainforest assembly have come from floristic elements which were carried on the
Indian Plate The Indian Plate (or India Plate) is a minor tectonic plate straddling the equator in the Eastern Hemisphere. Originally a part of the ancient continent of Gondwana, the Indian Plate broke away from the other fragments of Gondwana , began ...
and montane elements which have come from the
Australian Plate The Australian Plate is a major tectonic plate in the eastern and, largely, southern hemispheres. Originally a part of the ancient continent of Gondwana, Australia remained connected to India and Antarctica until approximately when India broke ...
(
Sahul __NOTOC__ Sahul (), also called Sahul-land, Meganesia, Papualand and Greater Australia, was a paleocontinent that encompassed the modern-day landmasses of mainland Australia, Tasmania, New Guinea, and the Aru Islands. Sahul was in the south- ...
). The Sahul component is now understood to include substantial two-way exchanges with
Sunda Sunda may refer to: Europe * Sunda, Faroe Islands India * Sunda (asura), an asura brother of Upasunda * Sunda (clan), a clan (gotra) of Jats in Haryana and Rajasthan, India Southeast Asia * Sundanese (disambiguation) ** Sundanese people ...
inclusive of lowland taxa. Evidence for the relative contributions of the great Asiatic floristic interchanges (GAFIs) with India and Sahul, respectively, to the flora of Malesia comes from contemporary lineage distributions, the
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
record, time-calibrated
phylogenies A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
, functional traits, and the spatial structure of genetic diversity. Functional trait and biome conservatism are noted features of montane austral lineages from Sahul (e.g., diverse
Podocarpaceae Podocarpaceae is a large family of mainly Southern Hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Pr ...
), whereas the abundance and diversity of lowland lineages, including groups such as
Syzygium ''Syzygium'' () is a genus of flowering plants that belongs to the myrtle family, Myrtaceae. The genus comprises about 1200 species, and has a native range that extends from Africa and Madagascar through southern Asia east through the Pacific. ...
( Myrtaceae) and the Asian dipterocarps (Dipterocarpoideae), reflect a less well understood combination of dispersal, ecology, and adaptive radiations. Thus, Malesian rainforest assembly has been shaped by sharply contrasting evolutionary origins and biogeographic histories.


See also

*
Nusantara Nusantara most commonly refers to: *Nusantara (archipelago), an Old Javanese term which initially referred to the conquered territories of the Majapahit empire, corresponding to present-day Indonesia *Nusantara (planned city), the future capital ci ...
* Phytochorion - floristic regions and provinces *
Coral Triangle The Coral Triangle (CT) is a roughly triangular area in the tropical waters around the Philippines, Indonesia, Malaysia, Papua New Guinea, the Solomon Islands and Timor-Leste. This area contains at least 500 species of reef-building corals in e ...
*
Wallacea Wallacea is a biogeographical designation for a group of mainly Indonesian islands separated by deep-water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as ...


References


External links

* * {{floristic regions Australasian realm Indomalayan realm Paleotropical flora Floristic regions