Maldives–Lakshadweep–Chagos Archipelago Tropical Moist Forests
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The Maldives–Lakshadweep–Chagos Archipelago tropical moist forests is a
tropical moist broadleaf forest Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat (ecology), habitat type defined by the World Wide Fund for Nature. Description TSMF is generally foun ...
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of l ...
in
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;;;;; ...
. It spans a chain of coralline islands in the Indian Ocean, including
Lakshadweep Lakshadweep (), also known as Laccadives (), is a union territory of India. It is an archipelago of 36 islands in the Arabian sea, located off the Malabar Coast. The name ''Lakshadweep'' means "one lakh islands" in Sanskrit, though the Lac ...
(Laccadive Islands), a union territory of
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
; the
Maldives Maldives (, ; dv, ދިވެހިރާއްޖެ, translit=Dhivehi Raajje, ), officially the Republic of Maldives ( dv, ދިވެހިރާއްޖޭގެ ޖުމްހޫރިއްޔާ, translit=Dhivehi Raajjeyge Jumhooriyyaa, label=none, ), is an archipelag ...
, an independent country; and the
British Indian Ocean Territory The British Indian Ocean Territory (BIOT) is an Overseas Territory of the United Kingdom situated in the Indian Ocean, halfway between Tanzania and Indonesia. The territory comprises the seven atolls of the Chagos Archipelago with over 1,000 ...
, an overseas territory of the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
.Wikramanayake, Eric; Eric Dinerstein; Colby J. Loucks; et al. (2002). ''Terrestrial Ecoregions of the Indo-Pacific: a Conservation Assessment.'' Washington, DC: Island Press.


Geography

The ecoregion is made up of low islands made mostly of coral sand, generally no more than 5 meters above sea level, and surrounded by extensive
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. Co ...
s. The islands rest on the
Chagos–Laccadive Ridge The Chagos–Laccadive Ridge (CLR), also known as the Chagos–Laccadive Plateau, is a prominent volcanic ridge and oceanic plateau extending between the Northern and the Central Indian Ocean. Extending from 10°S to 15°N, the CLR include ...
, an oceanic ridge and plateau, and extend 2,550 km north and south across the equator, from latitude 14º N to 8º S. Lakshadweep lies to the north, 300 km west of India's southwest coast. Lakshadweep consists of 36 small islands between 8º - 14º N, with a total land area of 32 km2. The Maldives are the largest island group, with approximately 1190 islands between 7º N and the equator. The Chagos Archipelago lies between 5° and 8° S, and constitute the British Indian Ocean Territory.
Diego Garcia Diego Garcia is an island of the British Indian Ocean Territory, a disputed overseas territory of the United Kingdom. It is a militarised atoll just south of the equator in the central Indian Ocean, and the largest of the 60 small islands o ...
, a huge atoll, is the largest of the Chagos islands. The coral sand that makes up the islands comes from the skeletons of reef corals which have been broken up into fine particles by wave action. Wind, waves, and currents form islands from the sand. The coral reefs provide the raw material for the islands, and the reefs provide protection from erosive waves and currents that allows the islands to form and persist. Coral sand is highly porous, and water and organic nutrients quickly leach through it. The islands' small size, low relief, and porous soil mean that, despite high rainfall, they have no running streams; rainfall percolates 1–3 meters below the ground to shallow freshwater
aquifer An aquifer is an underground layer of water-bearing, permeable rock, rock fractures, or unconsolidated materials (gravel, sand, or silt). Groundwater from aquifers can be extracted using a water well. Aquifers vary greatly in their characterist ...
s lying above salt water.


Climate

The climate of the island is tropical and equatorial, with warm temperatures that stay relatively constant throughout the year. Rainfall occurs mostly with the April-to-October
southwest monsoon A monsoon () is traditionally a seasonal reversing wind accompanied by corresponding changes in precipitation but is now used to describe seasonal changes in atmospheric circulation and precipitation associated with annual latitudinal oscill ...
, and there is a dry season from December to March associated with the northeast monsoon off the Asian continent. Rainfall varies from 1600 mm annually in dry parts of Lakshadweep to 3,800 mm per year in the southern Maldives.


Flora

The vegetation on the islands is dependent on the underlying soils. Pioneer plants colonize areas of fresh sand, and stabilize them. Areas with infertile soil are generally covered with sedges and shrubs that can tolerate drought and salt spray. Sedges of genus ''
Cyperus ''Cyperus'' is a large genus of about 700 species of sedges, distributed throughout all continents in both tropical and temperate regions. Description They are annual or perennial plants, mostly aquatic and growing in still or slow-moving w ...
'' and the shrub ''
Scaevola taccada ''Scaevola taccada'', also known as beach cabbage, sea lettuce, or beach naupaka, is a flowering plant in the family Goodeniaceae found in coastal locations in the tropical areas of the Indo-Pacific. It is a common beach shrub throughout the Arab ...
'' are the most common. In the Laccadive Islands the low tree ''
Argusia ''Tournefortia'', commonly known as soldierbush, is a genus of flowering plants in the borage family, Boraginaceae. It was first published under the name ''Pittonia'' by Charles Plumier in 1703, in honour of Joseph Pitton de Tournefort. Later, ...
'' is sometimes found with the sedges and shrubs. Where plant cover persists over time, organic material can accumulate in the soil and sustain larger and more diverse plants. Tropical rain forests grow where soils and fresh water are favorable. Most of the original forests have been cleared for coconut plantations and other crops, but small areas of native vegetation remain, some disturbed and others relatively intact. The Chagos Islands were the last settled and are the least disturbed. Plant communities on the islands include: * littoral shrubland is found along the seaward shores of the islands. ''Scaevola taccada'' is the dominant shrub species, growing from 2 to 5 meters high. The shrubs ''
Tournefortia argentea ''Heliotropium arboreum'' is a species of flowering plant in the borage family, Boraginaceae. It is native to tropical Asia including southern China, Madagascar, northern Australia, and most of the atolls and high islands of Micronesia and Polyn ...
'' and '' Suriana maritima'' are found close to the shore, along with the vine ''
Ipomoea macrantha ''Ipomoea'' () is the largest genus in the plant family Convolvulaceae, with over 600 species. It is a large and diverse group, with common names including morning glory, water convolvulus or water spinach, sweet potato, bindweed, moonfl ...
''. * ''Hernandia'' forest is dominated by the Chinese-lantern tree ''
Hernandia nymphaeifolia ''Hernandia nymphaeifolia'' is a species of plant in the Hernandiaceae family. Its common name is lantern tree. Description ''Hernandia nymphaeifolia'' is a tree with 5–22 m high. The leaves are narrowly or broadly ovate or subcircular. The 5- ...
'', sometimes with coconut palm (''Coco nucifera'') and ''
Terminalia catappa ''Terminalia catappa'' is a large tropical tree in the leadwood tree family, Combretaceae, native to Asia, Australia, the Pacific, Madagascar and Seychelles. Common names in English include country almond, Indian almond, Malabar almond, sea almon ...
'', and the smaller trees ''
Morinda citrifolia ''Morinda citrifolia'' is a fruit-bearing tree in the coffee family, Rubiaceae. Its native range extends across Southeast Asia and Australasia, and was spread across the Pacific by Polynesian sailors. The species is now cultivated throughout th ...
'', ''
Guettarda speciosa ''Guettarda speciosa'', with common names sea randa, or zebra wood, is a species of shrub in the family Rubiaceae found in coastal habitats in tropical areas around the Pacific Ocean, including the coastline of central and northern Queensland an ...
'', and '' Neisosperma oppositifolium''. The fern ''
Asplenium nidus :''See bird's-nest fern for other plants with this common name.'' ''Asplenium nidus'' is an epiphytic species of fern in the family Aspleniaceae, native to tropical southeastern Asia, eastern Australia, Hawaii (''ʻēkaha'' in Hawaiian), Poly ...
'' grows on the forest floor and as an
epiphyte An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...
in the trees. * ''Calophyllum'' forest is dominated by the tree ''
Calophyllum inophyllum ''Calophyllum inophyllum'' is a large evergreen plant, commonly called tamanu, oil-nut, mastwood, beach calophyllum or beautyleaf. It is native to tropical Asia and Wallacea. Due to its importance as a source of timber for the traditional shipbu ...
'', typically in pure stands, forming a dense canopy with little understory growth. The trees can grow to large size, with trunks up to two metres in diameter. * ''Barringtonia'' forest is dominated by ''
Barringtonia asiatica ''Barringtonia asiatica'' (fish poison tree, putat or sea poison tree) is a species of ''Barringtonia'' native to mangrove habitats from islands of the Indian Ocean in the west to tropical Asia and islands of the western Pacific Ocean. It is g ...
'', which can grow to a similar size as the ''Calophyllum'' trees. * ''Cordia'' forest is dominated by the tree ''
Cordia subcordata ''Cordia subcordata'' is a species of flowering tree in the borage family, Boraginaceae, that occurs in eastern Africa, South Asia, Southeast Asia, northern Australia and the Pacific Islands (including Hawaii). The plant is known by a variety o ...
'', either in pure stands or mixed with ''Hernandia nymphaeifolia'', ''Morinda citrifolia'', and ''Neisosperma oppositifolium''. * ''Pisonia'' forest occurs rarely, and is characterized by the tree ''
Pisonia grandis ''Pisonia grandis'', the grand devil's-claws, is a species of flowering tree in the ''Bougainvillea'' family, Nyctaginaceae. Description The tree has broad, thin leaves, smooth bark and bears clusters of green sweet-smelling flowers that matu ...
'', which can attain large size. The tree is associated with seabird colonies, since its sticky seeds are spread by birds and it can tolerate
guano Guano (Spanish from qu, wanu) is the accumulated excrement of seabirds or bats. As a manure, guano is a highly effective fertilizer due to the high content of nitrogen, phosphate, and potassium, all key nutrients essential for plant growth. G ...
-derived soils. * ''Guettarda'' forest is dominated by the low tree ''Guettarda speciosa'', with smaller numbers of ''Neisosperma oppositifolium''. It may be a transitional zone or stage between littoral shrubland and other forest types. * ''Casuarina'' woodland are woodlands dominated by ''
Casuarina equisetifolia ''Casuarina equisetifolia'', common names ''Coastal She-oak'' or ''Horsetail She-oak'' (sometimes referred to as the Australian pine tree or whistling pine tree outside Australia), is a she-oak species of the genus ''Casuarina''. The native ...
''. Its needle-like leaves typically form a thick layer of litter that limits the growth of understory plants, but sometimes it has an understory of shrubs including ''
Premna serratifolia ''Premna serratifolia'' is a small tree/shrub in the family Lamiaceae. It flowers and fruits between May and November.Described by Dr. N Sasidharan (Dr. B P Pal Fellow), Kerala Forest Research Institute, Peechi in India biodiversity portal http:/ ...
'' and ''Scaevola taccada''. * Coconut woodland includes both abandoned plantations and natural woodlands, either in pure stands or mixed with ''Hernandia, Guettarda'', and ''Neisosperma''. Understory plants include ''Asplenium nidus'', native herbs ''
Boerhavia albiflora ''Boerhavia'' is a genus of over 100 species in the Nyctaginaceae family. The genus was named for Herman Boerhaave, a Dutch Republic, Dutch botanist, and the genus name is frequently misspelled "''Boerhaavia''". Common names include spiderlings ...
,
Fimbristylis cymosa ''Fimbristylis cymosa'', commonly known as tropical fimbry, or St. John's sedge, is a sedge of the family Cyperaceae that is native to Australia. The rhizome, rhizomatous perennial grass-like or herb sedge typically grows to a height of . It blo ...
, and Stenotaphrum micranthum''. Freshwater marshes are found where the water table is high, and saltwater wetlands and
mangrove A mangrove is a shrub or tree that grows in coastal saline water, saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evoluti ...
forests grow in sheltered shoreline areas. A 2,500 km2 mangrove forest remains on
Minicoy Minicoy, locally known as Maliku (), is an island in Lakshadweep, India. Along with Viringili, it is on ''Maliku atoll'', the southernmost atoll of Lakshadweep archipelago. Administratively, it is a census town in the Indian union territory ...
island, and ''
Bruguiera parviflora ''Bruguiera parviflora'' is a tree in the family Rhizophoraceae. The specific epithet ' is from the Latin meaning "small flowers". Description ''Bruguiera parviflora'' grows up to tall with a trunk diameter of up to . The bark is pale grey to p ...
'' is the predominant mangrove species. The islands have few endemic plants, and the native plants are mostly typical of
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
coral islands. The ancient origins of the islands' plant species include Sri Lanka (44%), Africa (28%), and
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms, and also a phytogeographical floristic region in the Paleotropical Kingdom. It has been given different definitions. The ...
(25%). The present flora of the islands includes both native species and species that were intentionally or unintentionally introduced by humans.Whistler, W. Arthur (1997)."Botanical Survey of Diego Garcia Chagos Archipelago, British Indian Ocean Territory". in ''Diego Garcia Natural Resource Management Plan'', Prepared by Belt Collins Hawaii for the United States Navy, April 1997.


Fauna

The islands have few terrestrial animals. The islands are home to subspecies of
Indian flying fox The Indian flying fox (''Pteropus medius'', formerly ''Pteropus giganteus''), also known as the greater Indian fruit bat, is a species of flying fox native to the Indian subcontinent. It is one of the largest bats in the world. It is of interest ...
(''Pteropus giganteus ariel'') and
small flying fox The small flying fox, island flying fox or variable flying fox (''Pteropus hypomelanus'') is a species of flying fox in the family Pteropodidae. It is found in Australia, Cambodia, Indonesia, Malaysia, the Maldives, Myanmar, Papua New Guinea, the ...
(''Pteropus hypomelanus maris''). Both subspecies are endemic to the ecoregion. The Maldivian pond heron (''Ardeola grayii phillipsi''), a subspecies of the
Indian pond heron The Indian pond heron or paddybird (''Ardeola grayii'') is a small heron. It is of Old World origins, breeding in southern Iran and east to the Indian subcontinent, Burma, and Sri Lanka. They are widespread and common but can be easily missed whe ...
, is endemic to the islands. Thirteen or fourteen seabird species nest in the Maldives, often on small islets. The Chagos and Lakshadweep also have large rookeries. These include the
white tern The white tern or common white tern (''Gygis alba'') is a small seabird found across the tropical oceans of the world. It is sometimes known as the fairy tern, although this name is potentially confusing as it is also the common name of '' Sternu ...
(''Gygis alba monte''),
lesser frigatebird The lesser frigatebird (''Fregata ariel'') is a seabird of the frigatebird family Fregatidae. At around 75 cm (30 in) in length, it is the smallest species of frigatebird. It occurs over tropical and subtropical waters across the Indian ...
(''Fregata ariel iredalei''),
black-naped tern The black-naped tern (''Sterna sumatrana'') is an oceanic tern mostly found in tropical and subtropical areas of the Pacific and Indian Oceans. It is rarely found inland. Description The tern is about 30 cm long with a wing length of 21 ...
(''Sterna sumatrana''),
bridled tern The bridled tern (''Onychoprion anaethetus'') is a seabird of the family Laridae. It is a bird of the tropical oceans. The scientific name is from Ancient Greek. The genus comes from ' meaning "claw" or "nail", and , meaning "saw". The specific ...
(''Onychoprion anaethetus''), and
greater crested tern The greater crested tern Retrieved 28 February 2012 (''Thalasseus bergii''), also called crested tern or swift tern, is a tern in the family Laridae that nests in dense colonies on coastlines and islands in the tropical and subtropical Old World ...
(''Thalasseus bergii''). The red-footed booby (''Sula sula'') has a large population in the Chagos islands. The islands' native reptiles include two geckos, ''
Hemidactylus frenatus The common house gecko (''Hemidactylus frenatus'') is a gecko native to South and Southeast Asia. It is also known as the Asian house gecko, Pacific house gecko, wall gecko, house lizard, tayoto, chipkali or moon lizard. Most geckos are noctur ...
'' and '' H. parvimaculatus'',Agarwal, Ishan & Jablonski, Daniel & Bauer, A.. (2019). The identity and probable origin of the Hemidactylus geckos of the Maldives. Herpetological Journal. 29. 230-236. 10.33256/hj29.4.230236. the
Oriental garden lizard The oriental garden lizard (''Calotes versicolor''), also called the eastern garden lizard, Indian garden lizard, common garden lizard, bloodsucker or changeable lizard, is an agamid lizard found widely distributed in indo-Malaya. It has also bee ...
(''Calotes versicolor'') and another
agamid Agamidae is a family of over 300 species of iguanian lizards indigenous to Africa, Asia, Australia, and a few in Southern Europe. Many species are commonly called dragons or dragon lizards. Overview Phylogenetically, they may be sister to the ...
lizard, the
white-spotted supple skink The white-spotted supple skink (''Riopa albopunctata'') is a species of diurnal, terrestrial, insectivorous skink found in parts of tropical Asia. This species was first described by John Edward Gray based on type specimen collected by T. C. Je ...
(''Lygosoma albopunctatum''), and two snakes, the Indian wolf snake (''Lycodon aulicus'') and
Brahminy blind snake ''Indotyphlops braminus'', commonly known as the brahminy blind snake and other names, is a non-venomous blind snake species found mostly in Africa and Asia, but has been introduced in many other parts of the world. They are completely fossoria ...
(''Indotyphlops braminus''). Two native amphibians have been found on the islands, the
Indian burrowing frog The Indian burrowing frog (''Sphaerotheca breviceps'') is a species of frog found in South Asia (Bangladesh, Pakistan, Nepal, India, Sri Lanka and in Myanmar ). Description Its vomerine teeth form two oblique groups extending a little beyon ...
(''Sphaerotheca breviceps'') and
Asian common toad ''Duttaphrynus melanostictus'' is commonly called Asian common toad, Asian black-spined toad, Asian toad, black-spectacled toad, common Sunda toad, and Javanese toad. It is probably a complex of more than one true toad species that is widely d ...
(''Duttaphrynus melanostictus'').


History

Lakshadweep and the Maldives have been settled for thousands of years. The people of the Laccadives speak a dialect of
Malayalam Malayalam (; , ) is a Dravidian language spoken in the Indian state of Kerala and the union territories of Lakshadweep and Puducherry (Mahé district) by the Malayali people. It is one of 22 scheduled languages of India. Malayalam was des ...
, the predominant language in the adjacent Indian state of
Kerala Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South ...
. The people of the Maldives and Minicoy in Lakshadweep speak Maldivian, also known as Dhivehi, an
Indo-Aryan language The Indo-Aryan languages (or sometimes Indic languages) are a branch of the Indo-Iranian languages in the Indo-European language family. As of the early 21st century, they have more than 800 million speakers, primarily concentrated in India, Pa ...
most closely related to Sinhala spoken in nearby
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
. There is no record of permanent settlement on the Chagos Archipelago until 1793, when the French established a settlement and coconut plantations on Diego Garcia. The French brought enslaved people to the islands to work in the plantations. Their descendants, the
Chagossians The Chagossians (also Îlois or Chagos Islanders) are a currently exiled Creole ethnic group native to the Chagos Islands, specifically Diego Garcia, Peros Banhos, and the Salomon island chain, as well as other parts of the Chagos Archipelago ...
, speak a version of
Bourbonnais Creole Bourbonnais Creole is the group of French-based creole languages spoken in the western Indian Ocean. The close relation of the languages is from the similar historical and cultural backgrounds of the islands. The name is derived from the former ...
, a French-based creole language also spoken on
Réunion Réunion (; french: La Réunion, ; previously ''Île Bourbon''; rcf, label= Reunionese Creole, La Rényon) is an island in the Indian Ocean that is an overseas department and region of France. It is located approximately east of the island ...
and
Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label=Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It incl ...
in the southwestern Indian Ocean. The British consolidated control over the islands at the end of the 18th century. Lakshadweep was administered as part of
British India The provinces of India, earlier presidencies of British India and still earlier, presidency towns, were the administrative divisions of British governance on the Indian subcontinent. Collectively, they have been called British India. In one ...
, the Chagos Islands were administered from
Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label=Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It incl ...
, and the Maldives had local rulers under a British protectorate. Settlers brought food crops to the islands, and domestic animals including cats, chickens, goats, rabbits, house mice,
black rat The black rat (''Rattus rattus''), also known as the roof rat, ship rat, or house rat, is a common long-tailed rodent of the stereotypical rat genus ''Rattus'', in the subfamily Murinae. It likely originated in the Indian subcontinent, but is n ...
s (''Rattus rattus''),
Asian house shrew The Asian house shrew (''Suncus murinus'') is a shrew species native to South and Southeast Asia that has been listed as Least Concern on the IUCN Red List since 2008 because of its large population and wide distribution. It has been introduced i ...
s (''Suncus murinus'') and donkeys. Coconut plantations were expanded during the 19th century and became the islands' major export crop. Important island crops currently include bananas, coconuts, sweet potatoes, mangoes (''Mangifera indica''), chico (''Pouteria sapota''), yams, taro, millet, watermelons, citrus, and pineapples. India, including Lakshadweep, became independent in 1947. The Maldives became independent in 1965. The UK split administration of the Chagos Archipelago from that of Mauritius in 1965, three years prior to Mauritian independence. In the late 1960s, the UK government leased Diego Garcia to the United States, which turned it into a naval base. The Chagossians were expelled from the islands and resettled in Mauritius, the
Seychelles Seychelles (, ; ), officially the Republic of Seychelles (french: link=no, République des Seychelles; Creole: ''La Repiblik Sesel''), is an archipelagic state consisting of 115 islands in the Indian Ocean. Its capital and largest city, V ...
, and the UK. Mauritius still contests the UK's sovereignty over the islands, and the Chagossians continue to contest the legality of their expulsion and are seeking to return home.


Protected areas

A 2017 assessment found that 73 km2, or 26%, of the ecoregion is in protected areas.


External links

*


References

{{DEFAULTSORT:Maldives-Lakshadweep-Chagos Archipelago tropical moist forests Chagos Archipelago Ecoregions of Asia Ecoregions of India Environment of the Maldives Geography of the Maldives Indomalayan ecoregions Tropical and subtropical moist broadleaf forests