Macaca Nemestrina
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The southern pig-tailed macaque (''Macaca nemestrina''), also known as the
Sundaland Sundaland (also called Sundaica or the Sundaic region) is a biogeographical region of South-eastern Asia corresponding to a larger landmass that was exposed throughout the last 2.6 million years during periods when sea levels were lower. It ...
pig-tailed macaque and Sunda pig-tailed macaque, is a medium-sized
macaque The macaques () constitute a genus (''Macaca'') of gregarious Old World monkeys of the subfamily Cercopithecinae. The 23 species of macaques inhabit ranges throughout Asia, North Africa, and (in one instance) Gibraltar. Macaques are principally ...
that lives in
southern Thailand Southern Thailand, Southern Siam or Tambralinga is a southernmost cultural region of Thailand, separated from Central Thailand region by the Kra Isthmus. Geography Southern Thailand is on the Malay Peninsula, with an area of around , bounded ...
,
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
, and
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
. It is known locally as berok.


Etymology and taxonomy

The species
epithet An epithet (, ), also byname, is a descriptive term (word or phrase) known for accompanying or occurring in place of a name and having entered common usage. It has various shades of meaning when applied to seemingly real or fictitious people, di ...
, ''nemestrina'', is an adjective (derived from Latin ''Nemestrinus'', meaning "the god of groves") modified to agree in gender with the feminine
generic Generic or generics may refer to: In business * Generic term, a common name used for a range or class of similar things not protected by trademark * Generic brand, a brand for a product that does not have an associated brand or trademark, other ...
name. ''M. nemestrina'' formerly included the northern pig-tailed, Pagai Island, and Siberut macaques as
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
. All four are now considered separate species.


Description

As with other ''Macaca'' species, males are larger than females; while males are measured at in length and in weight, females are measured at in length and in weight. This macaque has buff-brown fur, with a darker dorsal area and lighter ventral area. Its common name refers to the short tail held semi-erect, resembling the tail of a pig.


Behaviour and ecology

''M. nemestrina'' is mainly
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
, but also a skilled climber. Unlike almost all primates, these macaques love water. They live in large groups that split into smaller groups during
daytime Daytime as observed on Earth is the period of the day during which a given location experiences natural illumination from direct sunlight. Daytime occurs when the Sun appears above the local horizon, that is, anywhere on the globe's hemis ...
when they are
foraging Foraging is searching for wild food resources. It affects an animal's Fitness (biology), fitness because it plays an important role in an animal's ability to survive and reproduce. Optimal foraging theory, Foraging theory is a branch of behaviora ...
. They are
omnivorous An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutri ...
, feeding mainly on fruits, seeds, berries,
cereal A cereal is any Poaceae, grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, Cereal germ, germ, and bran. Cereal Grain, grain crops are grown in greater quantit ...
s,
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
, and
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s. A study in peninsular
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
found them to be the primary, and perhaps the only, seed dispersers of the
rattan Rattan, also spelled ratan, is the name for roughly 600 species of Old World climbing palms belonging to subfamily Calamoideae. The greatest diversity of rattan palm species and genera are in the closed-canopy old-growth tropical forests of ...
species ''
Daemonorops ''Daemonorops'' was a genus of rattan palms in the family Arecaceae. Its species are now included within the genus ''Calamus''. Species found primarily in the tropics and subtropics of southeastern Asia with a few species extending into souther ...
calicarpa'' and ''
Calamus Calamus may refer to: Botany and zoology * ''Calamus'' (fish), a genus of fish in the family Sparidae * ''Calamus'' (palm), a genus of rattan palms * Calamus, the hollow shaft of a feather, also known as the quill * '' Acorus calamus'', the swe ...
castaneus''. There is a
hierarchy A hierarchy (from Greek: , from , 'president of sacred rites') is an arrangement of items (objects, names, values, categories, etc.) that are represented as being "above", "below", or "at the same level as" one another. Hierarchy is an important ...
among males, based on strength, and among females, based on
heredity Heredity, also called inheritance or biological inheritance, is the passing on of traits from parents to their offspring; either through asexual reproduction or sexual reproduction, the offspring cells or organisms acquire the genetic inform ...
. Thus, the daughter of the
alpha Alpha (uppercase , lowercase ; grc, ἄλφα, ''álpha'', or ell, άλφα, álfa) is the first letter of the Greek alphabet. In the system of Greek numerals, it has a value of one. Alpha is derived from the Phoenician letter aleph , whic ...
female will immediately be placed above all other females in the group. The alpha female leads the group, while the male role is more to manage conflict within the group and to defend it. Female
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregna ...
lasts around 5.7 months. She will give birth to one infant every two years. Weaning occurs at 4–5 months.
Sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans it might be considered synonymous with adulthood, but here puberty is the name for the process of biological sexual maturation, while adulthood is based on cultural definitio ...
is reached at 3–5 years. In
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
, they have been trained for 400 years to harvest
coconut The coconut tree (''Cocos nucifera'') is a member of the palm tree family ( Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or the ...
s.


Habitat and distribution

This
macaque The macaques () constitute a genus (''Macaca'') of gregarious Old World monkeys of the subfamily Cercopithecinae. The 23 species of macaques inhabit ranges throughout Asia, North Africa, and (in one instance) Gibraltar. Macaques are principally ...
is mostly found in rainforest up to 2000 meters, but will also enter
plantation A plantation is an agricultural estate, generally centered on a plantation house, meant for farming that specializes in cash crops, usually mainly planted with a single crop, with perhaps ancillary areas for vegetables for eating and so on. The ...
s and gardens.Payne, J., and C. M. Francis. 1998. ''A Field Guide to the Mammals of Borneo.'' The Sabah Society, Kota Kinabalu, Sabah. It is found in the southern half of the
Malay Peninsula The Malay Peninsula (Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area ...
(only just extending into southernmost
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
),
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...
,
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
and
Bangka Island Bangka is an island lying east of Sumatra, Indonesia. It is administered under the province of the Bangka Belitung Islands, being one of its namesakes alongside the smaller island of Belitung across the Gaspar Strait. The 9th largest island in In ...
. There are reports of the species having been present in
Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, borde ...
before 1950, but these were likely escaped pets. The only pig-tailed macaques in Singapore today are introduced monkeys.


References

*Maestripieri D, 1999. Changes in Social Behavior and Their Hormonal Correlates during Pregnancy in Pig-tailed Macaques. International Journal of Primatology 20 : 707-718. *Rodman PS, 1991. Structural differentiation of microhabitats of sympatricmacaca fascicularis andM. nemestrina in East Kalimantan, Indonesia. International Journal of Primatology 12 : 357-375. *Oi T, 1990. Patterns of dominance and affiliation in wild pig-tailed macaques (Macaca nemestrina nemestrina) in West Sumatra. International Journal of Primatology 11 : 339-356.


External links

* ARKive
images and movies of the Sunda pig-tailed macaque ''(Macaca nemestrina)''Biolib
{{DEFAULTSORT:macaque, pig-tailed, southern
southern pig-tailed macaque The southern pig-tailed macaque (''Macaca nemestrina''), also known as the Sundaland pig-tailed macaque and Sunda pig-tailed macaque, is a medium-sized macaque that lives in southern Thailand, Malaysia, and Indonesia. It is known locally as bero ...
Primates of Southeast Asia
southern pig-tailed macaque The southern pig-tailed macaque (''Macaca nemestrina''), also known as the Sundaland pig-tailed macaque and Sunda pig-tailed macaque, is a medium-sized macaque that lives in southern Thailand, Malaysia, and Indonesia. It is known locally as bero ...
Taxa named by Carl Linnaeus