Lake Malawi

Lake Malawi, also known as Lake Nyasa in Tanzania and Lago Niassa in Mozambique, is an African Great Lake and the southernmost lake in the East African Rift system, located between Malawi, Mozambique and Tanzania.

It is the fifth largest fresh water lake in the world by volume, the ninth largest lake in the world by area—and the third largest and second deepest lake in Africa. Lake Malawi is home to more species of fish than any other lake in the world, including at least 700 species of cichlids.Turner, Seehausen, Knight, Allender, and Robinson (2001). "How many species of cichlid fishes are there in African lakes?" ''Molecular Ecology'' 10: 793–806. The Mozambique portion of the lake was officially declared a reserve by the Government of Mozambique on June 10, 2011,WWF (10 June 2011)
"Mozambique’s Lake Niassa declared reserve and Ramsar site"
Retrieved 17 July 2014. while in Malawi a portion of the lake is included in Lake Malawi National Park.

Lake Malawi is a meromictic lake, meaning that its water layers do not mix. The permanent stratification of Lake Malawi's water and the oxic-anoxic boundary (relating to oxygen in the water) are maintained by moderately small chemical and thermal gradients.

Geography

Lake Malawi is between and long, and about wide at its widest point. The lake has a total surface area of about . The lake is at its deepest point, located in a major depression in the north-central part. Another smaller depression in the far north reaches a depth of . The southern half of the lake is shallower; less than in the south-central part and less than in the far south. The lake has shorelines on western Mozambique, eastern Malawi, and southern Tanzania. The largest river flowing into it is the Ruhuhu River, and there is an outlet at its southern end, the Shire River, a tributary that flows into the very large Zambezi River in Mozambique. Evaporation accounts for more than 80% of the water loss from the lake, considerably more than the outflowing Shire River.Park, L.E.; and A.S. Cohen (2011). Paleoecological response of ostracods to early Late Pleistocene lake-level changes in Lake Malawi, East Africa. Palaeogeography, Palaeoclimatology, Palaeoecology 303: 71–80. The outflows from Lake Malawi into the Shire River are vital for the economy as the water resources support hydropower, irrigation and downstream biodiversity. Concerns have been raised over the future climate change impacts of Lake Malawi due to the recent decline in lake levels and the overall drying trend. The climate in the lake region is already experiencing changes, with the temperatures predicted to increase throughout the country.

The lake is about southeast of Lake Tanganyika, another of the great lakes of the East African Rift.

The Lake Malawi National Park is located at the southern end of the lake.

File:Lake Malawi00.jpg, Lake Malawi (1967)
File:Mwaya Beach, Malawi.jpg, Mwaya Beach
File:Monoxylon beach Lake Malawi 1557.jpg, Beach at Cape Maclear near Monkey Bay

Geological history

Malawi is one of the major Rift Valley lakes and an ancient lake. The lake lies in a valley formed by the opening of the East African Rift, where the African tectonic plate is being split into two pieces. This is called a divergent plate tectonics boundary. Malawi has typically been estimated to be 1–2 million years old (mya),Givnish, T.J.; and K.J. Sytsma, editors (1997). Molecular Evolution and Adaptive Radiation, p. 598. . but more recent evidence points to a considerably older lake with a basin that started to form about 8.6 mya and deep-water condition first appeared 4.5 mya.Delvaux, D. (1995). Age of Lake Malawi (Nyasa) and water level fluctuations. Mus. roy. Afr. centr., Tervuren (Belg.), Dept. Geol. Min., Rapp. ann. 1993 & 1994: 99–108.Sturmbauer; Baric; Salzburger; Rüber; and Verheyen (2001). Lake Level Fluctuations Synchronize Genetic Divergences of Cichlid Fishes in African Lakes. Mol Biol Evol 18(2): 144–154.

The water levels have varied dramatically over time, ranging from almost below current levelCohen; Stone; Beuning; Park; Reinthal; Dettman; Scholz; Johnson; King; Talbot; Brown; and Ivory (2007). Ecological consequences of early Late Pleistocene megadroughts in tropical Africas. Proc Natl Acad Sci USA 104(42): 16422-16427. to above. During periods the lake dried out almost completely, leaving only one or two relatively small, highly alkaline and saline lakes in what currently are Malawi's deepest parts. A water chemistry resembling the current conditions only appeared about 60,000 years ago. Major low-water periods are estimated to have occurred about 1.6 to 1.0–0.57 million years ago (where it might have dried out completely), 420,000 to 250,000–110,000 years ago, about 25,000 years ago and 18,000–10,700 years ago. During the peak of the low-water period between 1390 and 1860 AD, it may have been below current water levels.

Water characteristics

The lake's water is alkaline ( pH 7.7–8.6) and warm with a typical surface temperature between , while deep sections typically are about .Stauffer, J.R.; and H. Madsen (2012). Schistosomiasis in Lake Malawi and the Potential Use of Indigenous Fish for Biological Control. Pp. 119–140 in: Rokni, M.B., editor. Schistosomiasis. . The thermocline is located at a depth of . The oxygen limit is at a depth of approximately , effectively restricting fish and other aerobic organisms to the upper part.Lowe-McConnell, R.H. (2003). Recent research in the African Great Lakes: Fisheries, biodiversity and cichlid evolution. Freshwater Forum 20(1): 4–64. The water is very clear for a lake and the visibility can be up to , but slightly less than half this figure is more common and it is below in muddy bays. Konings, Ad (1990). ''Ad Konings' Book of Cichlids and all the other Fishes of Lake Malawi.'' . However, during the rainy season months of January to March, the waters are more muddy due to muddy river inflows.

European discovery and colonisation

The Portuguese trader Candido José da Costa Cardoso was the first European to visit the lake in 1846. David Livingstone reached the lake in 1859, and named it Lake Nyasa. He also referred to it by a pair of nicknames: Lake of Stars and Lake of Storms. The Lake of Stars nickname came after Livingstone observed lights from the lanterns of the fishermen in Malawi on their boats, that resemble, from a distance, stars in the sky. Later, after experiencing the unpredictable and extremely violent gales that sweep through the area, he also referred to it as the Lake of Storms.

On 16 August 1914, Lake Malawi was the scene of a brief naval battle when the British gunboat , commanded by a Captain Rhoades, heard that World War I had broken out, and he received orders from the British Empire's high command to "sink, burn, or destroy" the German Empire's only gunboat on the lake, the ''Hermann von Wissmann'', commanded by a Captain Berndt. Rhoades's crew found the ''Hermann von Wissmann'' in a bay near Sphinxhaven, in German East African territorial waters. ''Gwendolen'' disabled the German boat with a single cannon shot from a range of about . This very brief gunboat conflict was hailed by ''The Times'' in England as the British Empire's first naval victory of World War I.

Borders

Tanzania–Malawi dispute

The partition of the lake's surface area between Malawi and Tanzania is under dispute. Tanzania claims that the international border runs through the middle of the lake. On the other hand, Malawi claims the whole of the surface of this lake that is not in Mozambique, including the waters that are next to the shoreline of Tanzania. Both sides cite the Heligoland Treaty of 1890 between Great Britain and Germany concerning the border. The wrangle in this dispute occurred when the British colonial government, just after they had captured Tanganyika from Germany, placed all of the waters of the lake under a single jurisdiction, that of the territory of Nyasaland, without a separate administration for the Tanganyikan portion of the surface. Later in colonial times, two jurisdictions were established.

The dispute came to a head in 1967 when Tanzania officially protested to Malawi; however nothing was settled. Occasional flare-ups of conflict occurred during the 1990s and in the 21st century. In 2012, Malawi's oil exploration initiative brought the issue to the fore, with Tanzania demanding that exploration cease until the dispute was settled.

Malawi–Mozambique border

In 1954, an agreement was signed between the British and the Portuguese making the middle of the lake their boundary with the exception of Chizumulu Island and Likoma Island, which were kept by the British and are now part of Malawi.

Transport

began service on the lake in 1901 as the SS ''Chauncy Maples'': a floating clinic and church for the Universities' Mission to Central Africa. She later served as a ferry and is currently being renovated into a mobile clinic at Monkey Bay. The renovation was expected to be complete during the first half of 2014, but was halted in 2017. entered service in 1935. The ferry entered service in 1951. In recent years she has often been out of service, but when operational she runs between Monkey Bay at the southern end of the lake to Karonga on the northern end, and occasionally to the Iringa Region of Tanzania. The ferry entered service in 1980. By 1982 she was carrying 100,000 passengers each year., but as of 2014 she was out of service. She normally serves the southern part of the lake but if ''Ilala'' was out of service she operated the route to Karonga. The Tanzanian ferry was built in 1988. Her operator was the Tanzania Railway Corporation Marine Division until 1997, when it became the Marine Services Company Limited. ''Songea'' plies weekly between Liuli and Nkhata Bay via Itungi and Mbamba Bay. The worst Lake disaster was a ship accident by the MV Vipya in 1946 which resulted in 145 deaths.

Wildlife

Wildlife found in and around Lake Malawi or Nyasa includes Nile crocodiles, hippopotamus, monkeys, and a significant population of African fish eagles that feed off fish from the lake.

Fish

Fishing

Lake Malawi has for millennia provided a major food source to the residents of its shores since its waters are rich in fish. Among the most popular are the four species of chambo, consisting of any one of four species in the subgenus ''Nyasalapia'' ('' Oreochromis karongae'', '' O. lidole'', '' O. saka'' and '' O. squamipinnis''), as well as the closely related '' O. shiranus''.Turner, G.F.; and N.C. Mwanyama (July 1992
Distribution and Biology of Chambo (Oreochromis spp.) in Lakes Malawi and Malombe.
Food and Agriculture Organization, Fisheries and Aquaculture Department, FI:DP/MLW/86/013, Field Document 21. Retrieved 13 April 2017. Other species that support important fisheries include the Lake Malawi sardine (''Engraulicypris sardella'') and the large kampango catfish (''Bagrus meridionalis''). Most fishing provides food for the increasing human population near the lake, but some are exported from Malawi. The wild population of fish is increasingly threatened by overfishing and water pollution. A drop in the lake's water level represents another threat, and is believed to be driven by water extraction by the increasing human population, climate change and deforestation. The chambo and kampango have been particularly overfished (the kampango declined by about 90% from 2006 to 2016, ''O. karongae'' and ''O. squamipinnis'' by about 94%, and ''O. lidole'' might already be extinct) and they are now seriously threatened. The IUCN recognises 117 species of Malawi cichlids as threatened; some of these have tiny ranges and may be restricted to rocky coastlines only a few hundred metres long.IUCN Red Lists
Geographic Patterns
. Eastern Africa. Retrieved 25 March 2017.

Cichlids

Lake Malawi is noted for being the site of evolutionary radiations among several groups of animals, most notably cichlid fish. There are at least 700 cichlid species in Lake Malawi, with some estimating that the actual figure is as high as 1,000 species. The actual number is labelled with some uncertainty because of the many undescribed species and the extreme variation among some species, making the task of delimiting them very complex. Except for four species ('' Astatotilapia calliptera'', '' Coptodon rendalli'', ''Oreochromis shiranus'' and ''Serranochromis robustus''), all cichlids in the lake are endemic to the Malawi system, which also includes nearby smaller Lake Malombe and the upper Shire River.Oliver, M.K. (12 April 2015)
The Tilapias of Lake Malawi.
MalawiCichlids. Retrieved 13 April 2017. Many of these have become popular among aquarium owners due to their bright colors. Recreating a Lake Malawi biotope to host cichlids became quite popular in the aquarium hobby. Most Malawi cichlids are found in relatively shallow coastal waters, but ''Diplotaxodon'' has been recorded down to depths of and several (especially ''Diplotaxodon'', ''Rhamphochromis'' and '' Copadichromis quadrimaculatus'') are known from pelagic waters.

The cichlids of the lake are divided into two groups and the vast majority of the species are haplochromines. The sister species to the Malawi haplochromines is ''Astatotilapia'' sp. Ruaha (a currently undescribed species from Great Ruaha River), and these two separated between 2.13 and 6.76 million years ago (mya).Genner; Ngatunga; Mzighani; Smith; and Turner (2015). Geographical ancestry of Lake Malawi’s cichlid fish diversity. Biol. Lett. 11: 2015023. The earliest divergence within the Malawi haplochromines occurred between 1.20 and 4.06 mya, but most radiations in this group are far younger; in extreme cases species may have diverged only a few hundred years ago. The Malawi haplochromines are mouthbrooders, but otherwise vary extensively in general behaviour and ecology. Within the Malawi haplochromines there are two main groups, the haps and the mbuna. The haps (they were formerly included in '' Haplochromis'') can be further subdivided into three subgroups: The relatively large, often more than long, and aggressive piscivores that roam various habitats in pursuit of prey, the open-water (although often not far from sand or rocks) utaka that feed in schools on zooplankton and typically are of medium size, and finally a subgroup of "aberrant" species that essentially are defined by them not fitting clearly into the other subgroups.Elieson, M
Haps Vs. Mbuna.
Retrieved 11 April 2017.Aquaticcommunity (2004–08
Haplochromis.
Retrieved 13 April 2017. Adult male haps generally display bright colors, while juveniles of both sexes and adult females typically show a silvery or grey coloration with sometimes irregular black bars or other markings. The second main haplochromine group are the mbuna, a name used both locally and popularly, which means "rockfish" in Tonga. They are found at rocky outcrops, territorially aggressive (although commonly found in high densities) and often specialised aufwuchs feeders. The mbuna species tend to be relatively small, mostly less than long, and often both sexes are brightly colored with males having egg-shaped yellow spots on their anal fin (a feature particularly prevalent in the mbuna, but not exclusive to this group).

The second group, the tilapia, comprises only six species in two genera in Lake Malawi: The redbreast tilapia (''Coptodon rendalli''), a widespread African species, is the only substrate-spawning cichlid in the lake. This large cichlid mainly feeds on macrophytes. The remaining are five mouthbrooding species of '' Oreochromis''; four chambo in the subgenus ''Nyasalapia'' (''O. karongae'', ''O. lidole'', ''O. saka'' and ''O. squamipinnis'') that are endemic to the Lake Malawi system, as well as the closely related ''O. shiranus'', which also is found in Lake Chilwa. The Malawi ''Oreochromis'' mainly feed on phytoplankton, reach lengths up to depending on the exact species, and are mostly black or silvery-gray with relatively indistinct dark bars. Male chambo have unique genital tassels when breeding, which aid in egg fertilisation in a manner comparable to the egg-spots on the anal fin of haplochromines.

Non-cichlids

The vast majority of the fish species in the lake are cichlids. Among the non-cichlid native fish are several species of cyprinids (in genera '' Barbus'', '' Labeo'' and '' Opsaridium'', and the Lake Malawi sardine ''Engraulicypris sardella''), airbreathing catfish ('' Bathyclarias'' and '' Clarias'', and the kampango ''Bagrus meridionalis''), mochokid catfish ('' Chiloglanis'' and Malawi squeaker ''Synodontis njassae''), '' Mastacembelus'' spiny eel, mormyrids ('' Marcusenius'', ''Mormyrops'' and '' Petrocephalus''), the African tetra '' Brycinus imberi'', the poeciliid '' Aplocheilichthys johnstoni'', the spotted killifish (''Nothobranchius orthonotus''), and the mottled eel (''Anguilla nebulosa'').

At a genus level, most of these are widespread in Africa, but ''Bathyclarias'' is entirely restricted to the lake.

Invertebrates

Molluscs

Lake Malawi is home to 28 species of freshwater snails (including 16 endemics) and 9 bivalves (2 endemics, '' Aspatharia subreniformis'' and the unionid '' Nyassunio nyassaensis''). The endemic freshwater snails are all members of the genera '' Bellamya'', '' Bulinus'', '' Gabbiella'', '' Lanistes'' and ''Melanoides''.

Lake Malawi is home to a total of four snail species in the genus ''Bulinus'', which is a known intermediate host of bilharzia. A survey in Monkey Bay in 1964 found two endemic species of snails of the genus ('' B. nyassanus'' and '' B. succinoides'') in the lake, and two non-endemic species ('' B. globosus'' and '' B. forskalli'') in lagoons separated from it. The latter species are known intermediate hosts of bilharzia, and larvae of the parasite were detected in water containing these, but in experiments C. Wright of the British Museum of Natural History was unable to infect the two species endemic to the lake with the parasites. The field workers, who spent many hours on and in the lake, did not find either ''B. globosus'' or ''B. forskalli'' in the lake itself. More recently, the disease has become a problem in the lake itself as the endemic ''B. nyassanus'' has become an intermediate host. This change, first noticed in the mid-1980s, is possibly related to a decline in snail-eating cichlids (for example, '' Trematocranus placodon'') due to overfishing and/or a new strain of the bilharzia parasite.

Crustaceans

Unlike Lake Tanganyika with its many endemic freshwater crabs and shrimp, there are few such species in Lake Malawi. The Malawi blue crab, '' Potamonautes lirrangensis'' ( syn. ''P. orbitospinus''), is the only crab in the lake and it is not endemic.Cumberlidge, N., and Meyer, K. S. (2011).
A revision of the freshwater crabs of Lake Kivu, East Africa.
' Journal Articles. Paper 30.Dobson, M. (2004).
Freshwater Crabs of Africa.
'' Freshwater Forum 21: 3–26. The atyid shrimp '' Caridina malawensis'' is endemic to the lake, but it is poorly known and has historically been confused with '' C. nilotica'', which is not found in the lake. Pelagic zooplanktonic species include two cladocerans (''Diaphanosoma excisum'' and ''Bosmina longirostris''), three copepods (''Tropodiaptomus cunningtoni'', '' Thermocyclops neglectus'' and '' Mesocyclops aequatorialis''),Darwall; Allison; Turner; and Irvine (2010). Lake of flies, or lake of fish? A trophic model of Lake Malawi. Ecological Modelling 221: 713–727. and several ostracods (including both described and undescribed species).

Lake flies

Lake Malawi is famous for the huge swarms of tiny, harmless lake flies, ''Chaoborus edulis''.Morris, B. (2004). Insects and Human Life, pp. 73–76. These swarms, typically appearing far out over water, can be mistaken for plumes of smoke and were also noticed by David Livingstone when he visited the lake.van Huis, A.; H. van Gurp; and M. Dicke (2012). The Insect Cookbook: Food for a Sustainable Planet, p. 31. Malawi Tourism:
Interesting seasonal highlights of Malawi.
Retrieved 8 April 2017. The aquatic larvae feed on zooplankton, spending the day at the bottom and the night in the upper water levels. When they pupate they float to the surface and transform into adult flies. The adults are very short-lived and the swarms, which can be several hundred metres tall and often have a spiraling shape, are part of their mating behaviour.Andrew, D. (30 June 2015)
What Are These Strange Looking "Clouds"?
IFLScience. Retrieved 8 April 2017. They lay their eggs at the water's surface and the adults die. The larvae are an important food source for fish, and the adult flies are important both to birds and local people, who collect them to make '' kungu'' cakes/burgers, a local delicacy with a very high protein content.

2015 mine leak

In January 2015, a sediment control tank collapsed at the Paladin Energy-owned uranium mine in Northern Malawi after a high intensity rain storm hit the area. It was revealed that approximately 50 litres of non radioactive material leaked into a local creek. Despite reports in local media of radioactive contamination the government conducted independent scientific tests on the local river system and found that there was no effect on the environment.

Swimming

The 25 km solo swim across Lake Malawi between Cape Ngomba and Senga Bay has been accomplished on 5 occasions by 16 swimmers

1992: Lewis Pugh 9hrs 52 minutes (UK/SA) and Otto Thanning (SA) 10hrs 5 minutes

2010: Abigail Brown (UK) 9hrs 45 minutes

2013: Milko van Gool (Netherlands) 8hrs 46 minutes and Kaitlin Harthoorn (US) 9hrs 17 minutes

2016: (current record) Jean Craven (SA), Robert Dunford (Kenya), Michiel Le Roux (SA), Samantha Whelpton (SA), Greig Bannatyne (SA), Haydn Von Maltitz (SA), Douglas Livingstone-Blevins (SA) 7hrs 53 mins

2019: Chris Stapley ( Eswatini) and Jay Azran (SA) 8hrs 40 minutes, Andrew Stevens (Australia) 10hrs 50 minutes, and Ruth Azran (SA) 11hrs 8 minutes.

In 2019, Martin Hobbs (SA), became the first person to swim the full length of Lake Malawi (54 days), as well as setting the world record for longest solo swim in a lake

See also

* 1989 Malawi earthquake
*2009 Karonga earthquakes
* Southeast Africa

References

Further reading

*
Recent study on Lake Malawi water levels reveals drought 100,000 years ago
*
*''Growing up in a Border District and Resolving the Tanzania-Malawi Lake Dispute: Compromise and concessions'', by Godfrey Mwakikagile, African Renaissance Press, 2022

{{DEFAULTSORT:Malawi

Ancient lakes
African Great Lakes
Lakes of the Great Rift Valley
Lakes of Malawi
Lakes of Tanzania
Malawi–Mozambique border
Malawi–Tanzania border crossings
International lakes of Africa
Ramsar sites in Mozambique
Lakes of Mozambique
Territorial disputes of Malawi
Territorial disputes of Tanzania