Luzon Tarictic Hornbill
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The Luzon hornbill (''Penelopides manillae''), sometimes called Luzon tarictic hornbill, is a species of
hornbill Hornbills (Bucerotidae) are a family (biology), family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a Casque (an ...
in the family
Bucerotidae Hornbills (Bucerotidae) are a family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a casque on the upper mandible ...
. It is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to forests on
Luzon Luzon (; ) is the largest and most populous island in the Philippines. Located in the northern portion of the Philippines archipelago, it is the economic and political center of the nation, being home to the country's capital city, Manila, as ...
and nearby islands in the northern
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
. As is the case with all Philippine
tarictic hornbill ''Penelopides'' is the genus of relatively small, primarily frugivorous hornbills restricted to forested areas of the Philippines. Their common name, tarictic hornbills, is an onomatopoetic reference to the main call of several of them. They have ...
s, it has been considered a
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of '' P. panini''.Kemp, A. C. (2001). Family Bucerotidae (Hornbills). Pp. 436-523 in: del Hoyo, J., Elliott, A., & Sargatal, J. eds. (2001). ''Handbook of the Birds of the World.'' Vol. 6. Mousebirds to Hornbills. Lynx Edicions, Barcelona.


Taxonomy

The Luzon hornbill was described by the French polymath
Georges-Louis Leclerc, Comte de Buffon Georges-Louis Leclerc, Comte de Buffon (; 7 September 1707 – 16 April 1788) was a French naturalist, mathematician, cosmologist, and encyclopédiste. His works influenced the next two generations of naturalists, including two prominent Fr ...
in 1780 in his ''Histoire Naturelle des Oiseaux''. The bird was also illustrated in a hand-coloured plate engraved by
François-Nicolas Martinet François-Nicolas Martinet (1731 - 1800) was a French engineer, engraver and naturalist. Martinet engraved the plates for numerous works on natural history, especially ornithology. Notable in particular are those for ''l'Ornithologia, sive Synop ...
in the ''Planches Enluminées D'Histoire Naturelle'' which was produced under the supervision of
Edme-Louis Daubenton Edme-Louis Daubenton (12 August 1730 – 12 December 1785) was a French naturalist. Daubenton was the cousin of another French naturalist, Louis Jean-Marie Daubenton. Georges-Louis Leclerc, the Comte de Buffon engaged Edme-Louis Daubenton to su ...
to accompany Buffon's text. Neither the plate caption nor Buffon's description included a scientific name but in 1783 the Dutch naturalist
Pieter Boddaert Pieter Boddaert (1730 – 6 May 1795) was a Dutch physician and natural history, naturalist. Early life, family and education Boddaert was the son of a Middelburg jurist and poet by the same name (1694–1760). The younger Pieter obtained his M.D ...
coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Buceros manillae'' in his catalogue of the ''Planches Enluminées''. The type locality is
Manila Manila ( , ; fil, Maynila, ), officially the City of Manila ( fil, Lungsod ng Maynila, ), is the capital of the Philippines, and its second-most populous city. It is highly urbanized and, as of 2019, was the world's most densely populate ...
in the Philippines. The Luzon hornbill is now placed in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Penelopides ''Penelopides'' is the genus of relatively small, primarily frugivorous hornbills restricted to forested areas of the Philippines. Their common name, tarictic hornbills, is an onomatopoetic reference to the main call of several of them. They ha ...
'' that was introduced in 1849 by the German naturalist
Ludwig Reichenbach Heinrich Gottlieb Ludwig Reichenbach (8 January 1793 – 17 March 1879) was a German botanist and ornithologist. It was he who first requested Leopold Blaschka to make a set of glass marine invertebrate models for scientific education and museu ...
in a plate of the hornbills. The origin of the generic name is uncertain but it may be a combination of the Latin ''pene'' meaning "almost" or "nearly", the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''lophos'' meaning "crest" and ''-oideēs'' "resembling". The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''manillae'' is a Latinized form of "Manila". There are two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
: the relatively widespread
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, and ''subniger'' from the islands of Polillo and
Patnanongan Patnanungan, officially the Municipality of Patnanungan ( tgl, Bayan ng Patnanungan), is a 5th class municipality in the province of Quezon, Philippines. According to the 2020 census, it has a population of 15,052 people. A shoe-shaped island m ...
. It was hunted for meat, but is now considered to be safe, as a new hunting ban has been enforced.


In captivity

This species is kept in captivity.
Jurong Bird Park Jurong Bird Park is an aviary and tourist attraction in Jurong, Singapore. The largest such bird park in Asia, it covers an area of on the western slope of Jurong Hill, the highest point in the Jurong region. It is one of the parks managed by ...
keeps some specimens in their hornbill complex. There are two pairs in two different aviaries and Jurong bred the species on one occasion.
Safari World Safari World is a tourist attraction in Bangkok, Thailand that consists of two parks named Marine Park and Safari Park, operated by Safari World Public Limited. The park was opened in 1988 with a total area of for its open zoo and for its bird ...
in
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
also has some Luzon hornbills, and several zoos in the Philippines have achieved some success in breeding them.
San Diego Wild Animal Park The San Diego Zoo Safari Park, originally named the San Diego Wild Animal Park until 2010, is an 1,800 acre (730 ha) zoo in the San Pasqual Valley area of San Diego, California, near Escondido. It is one of the largest tourist attractions in Sa ...
used to keep and breed the species, but now the only ones outside Asia are kept at zoos in Berlin and
Frankfurt Frankfurt, officially Frankfurt am Main (; Hessian: , "Frank ford on the Main"), is the most populous city in the German state of Hesse. Its 791,000 inhabitants as of 2022 make it the fifth-most populous city in Germany. Located on its na ...
. Berlin keeps the nominate subspecies and Frankfurt have a pair of the ''subniger''. There have been problems with tarictic hornbills in captivity because all species were previously considered
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organ ...
. For example, the birds at Frankfurt are listed as ''subniger'' but do not have the correct bill colouration.


Gallery

image:Polillo-Tariktik-Hornvogel 060819.jpg, Male Image:Polillo Tariktik-Hornvogel 2007-05-25 088.jpg, Male Image:Polillo Tariktik-Hornvogel 2007-05-25 086.jpg, Male File:Penelopides manillae -Luzon Tarictic Hornbill-8.jpg, Female File:Penelopides manilae 4.jpg, Skeleton


References

{{Taxonbar, from=Q1061873
Luzon hornbill The Luzon hornbill (''Penelopides manillae''), sometimes called Luzon tarictic hornbill, is a species of hornbill in the family Bucerotidae. It is endemic to forests on Luzon and nearby islands in the northern Philippines. As is the case with al ...
Birds of Luzon
Luzon hornbill The Luzon hornbill (''Penelopides manillae''), sometimes called Luzon tarictic hornbill, is a species of hornbill in the family Bucerotidae. It is endemic to forests on Luzon and nearby islands in the northern Philippines. As is the case with al ...
Taxonomy articles created by Polbot