''Lusovenator'' (meaning "Portuguese hunter") is a genus of

carcharodontosaurian Allosauroidea is a superfamily or clade of theropod dinosaurs which contains four families — the Metriacanthosauridae, Allosauridae, Carcharodontosauridae, and Neovenatoridae. Allosauroids, alongside the family Megalosauroidea, were among th ...

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

, from the

Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...

(

Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...

) Praia de Amoreira Porto-Novo Member and the Late Jurassic-

Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pro ...

(

Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...

Berriasian In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ago) ...

) Assenta Member of the

Lourinhã Formation The Lourinhã Formation () is a fossil rich geological formation in western Portugal, named for the municipality of Lourinhã. The formation is mostly Late Jurassic in age (Kimmeridgian/Tithonian), with the top of the formation extending into the ...

in present-day

Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...

. It includes one species, ''Lusovenator santosi''.

Discovery and naming

The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

was discovered during the 1980s by José Joaquim dos Santos and he donated his fossil collection to the Sociedade de História Natural around thirty years later. The unnamed holotype was described and placed in the

Allosauroidea Allosauroidea is a superfamily or clade of theropod dinosaurs which contains four family (biology), families — the Metriacanthosauridae, Allosauridae, Carcharodontosauridae, and Neovenatoridae. Allosauroids, alongside the family Megalosauroid ...

in 2017. The neotype was described in 2019 and both specimens were placed within

Carcharodontosauria Allosauroidea is a superfamily or clade of theropod dinosaurs which contains four families — the Metriacanthosauridae, Allosauridae, Carcharodontosauridae, and Neovenatoridae. Allosauroids, alongside the family Megalosauroidea, were among t ...

.Malafaia, E., P. Mocho, F. Escaso, P. Dantas, and F. Ortega. 2019. "Carcharodontosaurian remains (Dinosauria, Theropoda) from the Upper Jurassic of Portugal". Journal of Paleontology 93 : 157–172 The species ''Lusovenator santosi'' was named and described in 2020. The known remains consist of the holotype SHN.036 - "a partial postcranial skeleton preserving the odontoid, the atlantal intercentrum, a cervical vertebra, isolated cervical neural spines, dorsal vertebrae, fragments of sacral vertebrae, caudal vertebrae, chevrons, fragments of cervical and dorsal ribs, the right ilium, both pubes, and ischia", thought to represent a juvenile individual - and the neotype SHN.019, "a partial skeleton represented by a series of articulated caudal vertebrae and an almost complete right pes".

Etymology

The generic name ''Lusovenator'' is derived from

Lusitania Lusitania (; ) was an ancient Iberian Roman province located where modern Portugal (south of the Douro river) and a portion of western Spain (the present Extremadura and the province of Salamanca) lie. It was named after the Lusitani or Lusita ...

, the province of the

Roman Empire The Roman Empire ( la, Imperium Romanum ; grc-gre, Βασιλεία τῶν Ῥωμαίων, Basileía tôn Rhōmaíōn) was the post-Republican period of ancient Rome. As a polity, it included large territorial holdings around the Mediterr ...

that roughly matches present-day Portugal, affixed with the Latin ''venator'', meaning "hunter". The specific epithet, ''santosi'', refers to José Joaquim dos Santos, who discovered the remains.

Classification

The describers ran a phylogenetic analysis and recovered ''Lusovenator'' within the clade Carcharodontosauria, outside

Carcharodontosauridae Carcharodontosauridae (carcharodontosaurids; from the Greek καρχαροδοντόσαυρος, ''carcharodontósauros'': "shark-toothed lizards") is a group of carnivorous theropod dinosaurs. In 1931, Ernst Stromer named Carcharodontosauridae ...

and

Neovenatoridae Neovenatoridae is a proposed clade of carcharodontosaurian dinosaurs uniting some primitive members of the group such as ''Neovenator'' with the Megaraptora, a group of theropods with controversial affinities. Other studies recover megaraptorans ...

. Their

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

is shown below:

References

{{Taxonbar, from=Q97149714 Carcharodontosauria Kimmeridgian life Late Jurassic dinosaurs of Europe Jurassic Portugal Fossils of Portugal Lourinhã Formation Fossil taxa described in 2020 Tithonian life