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The Luquillo Experimental Forest (''Bosque experimental de Luquillo'') is a protected area of
tropical rainforest Tropical rainforests are rainforests that occur in areas of tropical rainforest climate in which there is no dry season – all months have an average precipitation of at least 60 mm – and may also be referred to as ''lowland equatori ...
in northeastern
Puerto Rico Puerto Rico (; abbreviated PR; tnq, Boriken, ''Borinquen''), officially the Commonwealth of Puerto Rico ( es, link=yes, Estado Libre Asociado de Puerto Rico, lit=Free Associated State of Puerto Rico), is a Caribbean island and Unincorporated ...
. The experimental forest is located in the
Sierra de Luquillo The Sierra de Luquillo is a mountain range located in the northeastern part of Puerto Rico. Also known as the Luquillo Mountains, these are steep-sided, densely-forested mountains rising to elevations of around , the highest point being the summit ...
some east of San Juan, the capital of the island. It is a
UNESCO Biosphere Reserve Man and the Biosphere Programme (MAB) is an intergovernmental scientific program, launched in 1971 by UNESCO, that aims to establish a scientific basis for the improvement of relationships between people and their environments. MAB's work engag ...
and is used for research into
silviculture Silviculture is the practice of controlling the growth, composition/structure, and quality of forests to meet values and needs, specifically timber production. The name comes from the Latin ('forest') and ('growing'). The study of forests and wo ...
, forest regeneration, and other purposes.


History

The Tropical Forest Research Station was founded in 1940 and became a center for ecosystem research. It was designated as a UNESCO Biosphere Reserve in 1976 with the objective of understanding "the long-term dynamics of tropical forest ecosystems characterized by large-scale, infrequent disturbance, rapid processing of organic material, and high habitat and species diversity". Although the experimental forest is located within
El Yunque National Forest El Yunque National Forest ( es, Bosque Nacional El Yunque), formerly known as the Caribbean National Forest (or ''Bosque Nacional del Caribe''), is a forest located in northeastern Puerto Rico. It is the only tropical rainforest in the United Sta ...
, the two have different objectives. In a forest management plan drawn up in 1956, approximately were to be used for commercial timber production, mostly the lower, flatter part of the site. The remaining , largely mountain peaks and steep slopes, were considered non-commercial and were set aside for research and other purposes. Much of the experimental area is used for silviculture and reforestation research, however certain areas have been left completely unmanaged, and are of global importance for use in long term ecological studies. The experimental forest is one of 26 sites run by the
Long Term Ecological Research Network The Long Term Ecological Research Network (LTER) consists of a group of over 1800 scientists and students studying ecological processes over extended temporal and spatial scales. Twenty-eight LTER sites cover a diverse set of ecosystems. It is par ...
, a group of international scientists studying
ecological processes Ecosystem ecology is the integrated study of living ( biotic) and non-living (abiotic) components of ecosystems and their interactions within an ecosystem framework. This science examines how ecosystems work and relates this to their components s ...
over long time scales. The facility was established in 1988 to study the "long-term effects of natural and human disturbances on tropical forests and streams in the Luquillo mountains". The aspects studied include how
hurricanes A tropical cyclone is a rapidly rotating storm system characterized by a low-pressure center, a closed low-level atmospheric circulation, strong winds, and a spiral arrangement of thunderstorms that produce heavy rain and squalls. Depend ...
and
droughts A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, an ...
have affected the environment, and the effects of changes in agricultural practices and increased urbanization. A central database is maintained, collaboration between scientists, students and volunteers is encouraged, and efforts are made to involve the local community.


Sierra de Luquillo

The
Sierra de Luquillo The Sierra de Luquillo is a mountain range located in the northeastern part of Puerto Rico. Also known as the Luquillo Mountains, these are steep-sided, densely-forested mountains rising to elevations of around , the highest point being the summit ...
or Luquillo mountains are about high with steep, convoluted slopes.
Trade winds The trade winds or easterlies are the permanent east-to-west prevailing winds that flow in the Earth's equatorial region. The trade winds blow mainly from the northeast in the Northern Hemisphere and from the southeast in the Southern Hemisph ...
off the
Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
cool as they rise over the mountains, bringing an annual rainfall of . The mountains are well-forested and are clad in mist for most of the year. The Luquillo Experimental Forest has an area of about and encompasses five different vegetation zones; montane wet forest, montane rainforest, wet forest, rainforest, and a small area of moist forest in the southwestern part. The experimental forest site is wholly inside the
El Yunque National Forest El Yunque National Forest ( es, Bosque Nacional El Yunque), formerly known as the Caribbean National Forest (or ''Bosque Nacional del Caribe''), is a forest located in northeastern Puerto Rico. It is the only tropical rainforest in the United Sta ...
, which covers a large part of the mountain range to the southwest of the city of
Luquillo Luquillo () is a town and municipality of Puerto Rico located in the northeast coast, northwest of Fajardo; and east of Rio Grande. Luquillo is spread over 5 barrios and Luquillo Pueblo (the downtown area and the administrative center of the ci ...
. The composition of the forest varies according to the elevation, soil types and other factors. Compared to most mainland tropical forests the site has a low floral biodiversity; for example, only eight species of tree account for 75% of tree density. The lower parts are dominated by the Tabonuco tree (''Dacryodes excelsa''), the lower middle slopes by the Palo colorado (''Cyrilla racemiflora''), and the upper middle is predominantly Palma de sierra (''Prestoea montana)''. The upper slopes, with their perpetual cloud cover and permanently saturated soils, are clad with trees of low stature and are known as
dwarf forest Dwarf forest, elfin forest, or pygmy forest is an uncommon ecosystem featuring miniature trees, inhabited by small species of fauna such as rodents and lizards. They are usually located at high elevations, under conditions of sufficient air humi ...
. The dominant tree species here in the upper elevations include: ''Tabebuia rigida'', ''Eugenia borinquensis'', and ''Calycogonium squamulosum''.


Research

Over 2,500 research studies having been performed in the Luquillo Experimental Forest. One area of research is the resilience of the ecosystem in the face of hurricanes, landslides, and such long term changes as alterations in agricultural practices and use. A study in 2018 compared the biomass of animals collected in the forest with data from the 1970s and found a large decline, which the authors attributed to
global warming In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
, although the conclusions were disputed by a larger group of researchers working in the area, who were unable to replicate most of the findings, including the temperature change.


References

{{Authority control Ecological experiments Protected areas of Puerto Rico National Forests of Puerto Rico