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Ludwig Heinrich Philipp Döderlein (3 March 1855, Bad Bergzabern – 23 April 1936,
Munich Munich ( ; german: München ; bar, Minga ) is the capital and most populous city of the States of Germany, German state of Bavaria. With a population of 1,558,395 inhabitants as of 31 July 2020, it is the List of cities in Germany by popu ...
) was a
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
. He specialized in
echinoderms An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea li ...
, particularly
sea star Starfish or sea stars are Star polygon, star-shaped echinoderms belonging to the class (biology), class Asteroidea (). Common usage frequently finds these names being also applied to brittle star, ophiuroids, which are correctly referred to ...
s,
sea urchin Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) of ...
s, and
crinoid Crinoids are marine animals that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or comatulids, which are ...
s. He was one of the first European zoologists to have the opportunity to do research work in
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
from 1879 to 1881. Today, he is considered one of the most important pioneers of marine biological research in Japan. He was the director and curator of the
Musée zoologique de la ville de Strasbourg The Musée zoologique de la ville de Strasbourg is a natural history museum displaying the zoological collections of the city of Strasbourg, managed and curated by the University of Strasbourg. The museum is closed since September 2019 and until ...
from 1882 to 1919. He headed the
Zoologische Staatssammlung München The Bavarian State Collection of Zoology (german: Zoologische Staatssammlung München) or ZSM is a major German research institution for zoological systematics in Munich. It has over 20 million zoological specimens. It is one of the largest natur ...
from 1923 to 1927 and was Professor of Zoology in the
Ludwig Maximilian University of Munich The Ludwig Maximilian University of Munich (simply University of Munich or LMU; german: Ludwig-Maximilians-Universität München) is a public research university in Munich, Germany. It is Germany's sixth-oldest university in continuous operatio ...
.


Biography

Ludwig Döderlein was born in Bad Bergzabern, then
Kingdom of Bavaria The Kingdom of Bavaria (german: Königreich Bayern; ; spelled ''Baiern'' until 1825) was a German state that succeeded the former Electorate of Bavaria in 1805 and continued to exist until 1918. With the unification of Germany into the German E ...
, on March 3, 1855. He went to school in
Bayreuth Bayreuth (, ; bar, Bareid) is a town in northern Bavaria, Germany, on the Red Main river in a valley between the Franconian Jura and the Fichtelgebirge Mountains. The town's roots date back to 1194. In the 21st century, it is the capital of U ...
from 1864 to 1873. From 1873 to 1875 he studied
natural science Natural science is one of the branches of science concerned with the description, understanding and prediction of natural phenomena, based on empirical evidence from observation and experimentation. Mechanisms such as peer review and repeatab ...
s in the
University of Erlangen A university () is an institution of higher (or tertiary) education and research which awards academic degrees in several academic disciplines. Universities typically offer both undergraduate and postgraduate programs. In the United States, th ...
, where he also worked as an assistant to the Zoologist
Emil Selenka Emil Selenka (27 February, 1842, Braunschweig – 20 February, 1902, Munich) was a German zoologist. He is known for his research on invertebrates and apes and the scientific expeditions he organized to Southeast Asia and South America. Lif ...
in the summer of 1875. From 1875 to 1876 he took two semesters at the
Ludwig Maximilian University of Munich The Ludwig Maximilian University of Munich (simply University of Munich or LMU; german: Ludwig-Maximilians-Universität München) is a public research university in Munich, Germany. It is Germany's sixth-oldest university in continuous operatio ...
. After graduating, Döderlein moved to the
University of Straßburg The University of Strasbourg (french: Université de Strasbourg, Unistra) is a public research university located in Strasbourg, Alsace, France, with over 52,000 students and 3,300 researchers. The French university traces its history to the e ...
. There he completed his
Doctorate of Philosophy A Doctor of Philosophy (PhD, Ph.D., or DPhil; Latin: or ') is the most common degree at the highest academic level awarded following a course of study. PhDs are awarded for programs across the whole breadth of academic fields. Because it is a ...
in Mathematics and Natural Sciences on June 26, 1877. From 1876 to 1878 he worked for four semesters in the zoology department of the University of Straßburg as an assistant to the German zoologist and phycologist
Eduard Oscar Schmidt Eduard Oscar Schmidt (21 February 1823, in Torgau – 17 January 1886, in Kappelrodeck) was a German zoologist and phycologist. Biography He initially studied mathematics and science at Halle, then continued his education in Berlin, where he ...
. The prospects for an academic career were poor, so for a time, he worked as a schoolteacher in the Alsatian town of
Mulhouse Mulhouse (; Alsatian language, Alsatian: or , ; ; meaning ''Mill (grinding), mill house'') is a city of the Haut-Rhin Departments of France, department, in the Grand Est Regions of France, region, eastern France, close to the France–Switzerl ...
. There he met and befriended the
Japanese Japanese may refer to: * Something from or related to Japan, an island country in East Asia * Japanese language, spoken mainly in Japan * Japanese people, the ethnic group that identifies with Japan through ancestry or culture ** Japanese diaspor ...
student Kenji Oosawa (1852–1927), who was studying medicine and physiology at the University of Strasbourg. Oosawa arranged for Döderlein to be invited to
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
to work as a Professor of Natural History in the Faculty of Medicine of the newly established
University of Tokyo , abbreviated as or UTokyo, is a public research university located in Bunkyō, Tokyo, Japan. Established in 1877, the university was the first Imperial University and is currently a Top Type university of the Top Global University Project by ...
. Döderlein accepted gratefully. Döderlein was one of the first European academics invited to Japan during the
Meiji Restoration The , referred to at the time as the , and also known as the Meiji Renovation, Revolution, Regeneration, Reform, or Renewal, was a political event that restored practical imperial rule to Japan in 1868 under Emperor Meiji. Although there were ...
era, when Japan was undergoing rapid modernization after the end of the isolation period (
Sakoku was the Isolationism, isolationist Foreign policy of Japan, foreign policy of the Japanese Tokugawa shogunate under which, for a period of 265 years during the Edo period (from 1603 to 1868), relations and trade between Japan and other countri ...
). He stayed from 1879 to 1881 as an ''oyatoi gaikokujin'' ("foreign employee") professor. During this time, he collected and preserved marine life extensively, particularly fish,
sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...
s,
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s,
crinoid Crinoids are marine animals that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or comatulids, which are ...
s, sea urchins,
cnidarian Cnidaria () is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter. Their distinguishing feature is cnidocytes, specialized cells that th ...
s, and bryozoans. In addition, he also occasionally collected specimens of
amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...
s (including a
giant salamander The Cryptobranchidae are a family of fully aquatic salamanders commonly known as the giant salamanders. They include the largest living amphibians. The family is native to China, Japan, and the eastern United States. They constitute one of two li ...
),
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s,
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s, and
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
s. At first, he obtained most of his specimens from the
fish market A fish market is a marketplace for selling fish and fish products. It can be dedicated to wholesale trade between fishermen and fish merchants, or to the sale of seafood to individual consumers, or to both. Retail fish markets, a type of wet mar ...
s and gift shops of
Tokyo Tokyo (; ja, 東京, , ), officially the Tokyo Metropolis ( ja, 東京都, label=none, ), is the capital and largest city of Japan. Formerly known as Edo, its metropolitan area () is the most populous in the world, with an estimated 37.468 ...
and the island of
Enoshima is a small offshore island, about in circumference, at the mouth of the Katase River which flows into the Sagami Bay of Kanagawa Prefecture, Japan. Administratively, Enoshima is part of the mainland city of Fujisawa, and is linked to ...
. Later, he himself fished for specimens directly from
Sagami Bay lies south of Kanagawa Prefecture in Honshu, central Japan, contained within the scope of the Miura Peninsula, in Kanagawa, to the east, the Izu Peninsula, in Shizuoka Prefecture, to the west, and the Shōnan coastline to the north, while the ...
. At the end of the two-year period, Döderlein returned to Europe. Despite the difficulties associated with it, he brought his extensive collection of more than 3,550 specimens of at least 372
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
back with him. After his return, Döderlein became the director and curator of
Musée zoologique de la ville de Strasbourg The Musée zoologique de la ville de Strasbourg is a natural history museum displaying the zoological collections of the city of Strasbourg, managed and curated by the University of Strasbourg. The museum is closed since September 2019 and until ...
between 1882 and 1919. There he devoted much of his life cataloging its rich collection of marine fauna from the
Far East The ''Far East'' was a European term to refer to the geographical regions that includes East and Southeast Asia as well as the Russian Far East to a lesser extent. South Asia is sometimes also included for economic and cultural reasons. The ter ...
. During his tenure, he hired specialists like Johannes Thiele,
Franz Eilhard Schulze Franz Eilhard Schulze (22 March 1840 – 2 November 1921) was a German anatomist and zoologist born in Eldena, near Greifswald. Biography He studied at the Universities of Bonn and Rostock. In 1863, he received his doctorate from Rostock, where ...
, and
Arnold Edward Ortmann Arnold Edward Ortmann (April 8, 1863 – January 3, 1927) was a Prussian-born United States naturalist and zoologist who specialized in malacology. Biography Ortmann was born in Magdeburg, Prussia on April 8, 1863. A student of Ernst Haeckel, ...
to study his Japanese collection. He also taught zoology at the University of Strasbourg during this period. At the turn of the century, he developed severe symptoms of
tuberculosis Tuberculosis (TB) is an infectious disease usually caused by '' Mycobacterium tuberculosis'' (MTB) bacteria. Tuberculosis generally affects the lungs, but it can also affect other parts of the body. Most infections show no symptoms, in ...
. As a result, he was denied a promotion to Professor of Zoology because he could no longer teach effectively. From January to May 1901, in an effort to cure his ailment, he travelled to
Biskra Biskra ( ar, بسكرة ; ; Latin Vescera) is the capital city of Biskra Province, Algeria. In 2007, its population was recorded as 307,987. Biskra is located in northeastern Algeria, about 248 miles (400 km) from Algiers, 71 miles (115&nbs ...
in
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
where he spent most of his time studying the local fauna. After the end of the
First World War World War I (28 July 1914 11 November 1918), often abbreviated as WWI, was one of the deadliest global conflicts in history. Belligerents included much of Europe, the Russian Empire, the United States, and the Ottoman Empire, with fightin ...
in 1919, Alsace was ceded back to
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
by
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
under the
Treaty of Versailles The Treaty of Versailles (french: Traité de Versailles; german: Versailler Vertrag, ) was the most important of the peace treaties of World War I. It ended the state of war between Germany and the Allied Powers. It was signed on 28 June ...
. Döderlein was removed from directorship of the Museum of Strasbourg and was deported back to Germany by the French government. His family were not allowed to take anything with them per the treaty, forcing him to leave all his private property as well as his Japanese zoological collection behind. To make matters worse, Germany (then under the
Weimar Republic The Weimar Republic (german: link=no, Weimarer Republik ), officially named the German Reich, was the government of Germany from 1918 to 1933, during which it was a constitutional federal republic for the first time in history; hence it is al ...
) was struggling from the post-war economic depression. He spent the rest of his life heading the
Zoologische Staatssammlung München The Bavarian State Collection of Zoology (german: Zoologische Staatssammlung München) or ZSM is a major German research institution for zoological systematics in Munich. It has over 20 million zoological specimens. It is one of the largest natur ...
(Bavarian State Collection of Zoology) and teaching zoology in the University of Munich. He made numerous attempts to recover his Japanese collection but despite strong international support (including from the Smithsonian) he had little success. He also attempted to visit Strasbourg personally but was again thwarted by the growing hostility between Germany and France prior to the
Second World War World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the vast majority of the world's countries—including all of the great powers—forming two opposin ...
. He died in Munich on April 23, 1936 at the age of 81.


Legacy

Döderlein is considered as one of the last "great naturalists". His published work were varied, including papers on
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
,
tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a pig, with a short, prehensile nose trunk. Tapirs inhabit jungle and forest regions of South and Central America, with one species inhabit ...
s,
elephant Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae an ...
s, and even
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
s. Most of his work, however, was on
marine biology Marine biology is the scientific study of the biology of marine life, organisms in the sea. Given that in biology many phyla, families and genera have some species that live in the sea and others that live on land, marine biology classifies s ...
, with a special interest in
echinoderm An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea ...
s. He was mostly forgotten during the mid-Twentieth century until the rediscovery of his extensive collections at the Musée zoologique de la ville de Strasbourg by the Japanese marine biologist Shunsuke Mawatari. From 1997 to 2005, a Japanese team documented his collections in Strasbourg and other museums in
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
, Germany, and
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
, revealing the true extent of his work. He is now recognized as one of the pioneers of natural history research in Japan. He also started the long tradition of focused research on the ecologically rich Sagami Bay. His observations on its fauna is believed to have been the primary reason for the establishment of the Misaki Marine Biological Station in Sagami Bay in 1884 by the Japanese zoologist
Kakichi Mitsukuri was a Japanese people, Japanese Zoology, zoologist. Biography Kakichi Mitsukuri was born in Edo. In 1873 he came to the United States, where he received a Doctor of Philosophy, Ph.D. degree from Yale University, Yale in 1879 and from Johns Hop ...
. Döderlein is commemorated in the scientific name of a species of Asian snake, '' Calamaria doederleini''Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . ("Doederlein", p. 73). and in the fish genus '' Doederleinia''.


See also

* :Taxa named by Ludwig Heinrich Philipp Döderlein


References


External links


Digitized works of Ludwig Döderlein
from the
Biodiversity Heritage Library The Biodiversity Heritage Library (BHL) is the world’s largest open access digital library for biodiversity literature and archives. BHL operates as worldwide consortiumof natural history, botanical, research, and national libraries working toge ...
{{DEFAULTSORT:Doderlein, Ludwig Heinrich Philipp Zoologists with author abbreviations 1855 births 1936 deaths 19th-century German zoologists 20th-century German zoologists Scientists from Munich People from Bad Bergzabern University of Strasbourg faculty University of Tokyo faculty Ludwig Maximilian University of Munich faculty Members of the Bavarian Academy of Sciences Members of the German Academy of Sciences Leopoldina Evolutionary biologists