The long-winged harrier (''Circus buffoni'') is a common
bird of prey
Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predat ...
species endemic to
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
. They are members of the family
Accipitridae
The Accipitridae is one of the three families within the order Accipitriformes, and is a family of small to large birds with strongly hooked bills and variable morphology based on diet. They feed on a range of prey items from insects to medium-s ...
, under the genus ''
Circus
A circus is a company of performers who put on diverse entertainment shows that may include clowns, acrobats, trained animals, trapeze acts, musicians, dancers, hoopers, tightrope walkers, jugglers, magicians, ventriloquists, and unicyclis ...
'' which encompasses other
harrier species. Its range encompasses most of
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
, in
grassland
A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
and
wetland
A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The p ...
all across the continent. The long-winged harrier is a carnivorous bird, and will feed on many different animals found in its habitat.
Like
owl
Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
s and other
harrier species, the long-winged harrier has a distinctive
facial disc
In ornithology, the facial disc is the concave collection of feathers on the face of some birds—most notably owls—surrounding the eyes. The concavity of the facial disc forms a circular paraboloid that collects sound waves and directs those ...
, which is used to triangulate the bird's hearing while it is hunting.
The long-winged harrier is classified as a species of least concern by the IUCN Red List, however their population has been observed to decline due to habitat degradation by human activity.
Taxonomy
The long-winged harrier was
formally described in 1788 by the German naturalist
Johann Friedrich Gmelin in his revised and expanded edition of
Carl Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his Nobility#Ennoblement, ennoblement in 1761 as Carl von Linné#Blunt, Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalise ...
's ''
Systema Naturae''. He placed it with the falcons, eagles and their relatives in the
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''
Falco'' and coined the
binomial name ''Falco buffoni''. Gmelin based his description on the "Cayenne ringtail" that had been described in 1781 by the English ornithologist
John Latham in his multi-volume work ''A General Synopsis of Birds''. Latham had examined a specimen from
Cayenne that formed part of a private collection in London. The specimen had been given the title "Duc de Buffon". The long-winged harrier is now one of 16 harriers placed in the genus ''
Circus
A circus is a company of performers who put on diverse entertainment shows that may include clowns, acrobats, trained animals, trapeze acts, musicians, dancers, hoopers, tightrope walkers, jugglers, magicians, ventriloquists, and unicyclis ...
'' that was introduced in 1799 by the French naturalist
Bernard Germain de Lacépède.
The genus name ''Circus'' is derived from the
Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
''kirkos'', referring to a bird of prey named for its circling flight (''kirkos'', "circle"), probably the
hen harrier
The hen harrier (''Circus cyaneus'') is a bird of prey. It breeds in Eurasia. The term "hen harrier" refers to its former habit of preying on free-ranging fowl.
It migrates to more southerly areas in winter. Eurasian birds move to southern Eur ...
. The specific epithet ''buffoni'' is from the name of the French naturalist, the
Comte de Buffon. The long-winged harrier does not have any recognised
subspecies.
[
]
Description
The long-winged is mid-sized bird of prey, and like most birds of prey species, the females are larger than the males. Males will weigh from and females will be slightly heavier, ranging from . Their length can range between and their wingspan ranges between .[ As other harrier species, the lon-winged harrier can be identified by its long and narrow tail and its especially long wings.] Their wings and their back are mostly grey, with the primary feathers
Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tai ...
on the wings being much darker than the rest of the wing feathers, which are streaked with variants of white, grey and brown.
The long-winged harrier can arbor 2 different morphs, a dark morph and a light morph. In dark morph individuals, the belly is all black with females having more of a brown undertone. The light morph individuals can be identified by their white underparts, which contrasts heavily with their dark morph counterpart. Juvenile long-winged harriers usually have the same colour patterns as females, but they have a more speckled chest. The long-winged harrier morphologically resembles its close relative, the cinereous harrier (''Circus cinereus''), which can also be found in South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
, but their range rarely overlaps.
As all other birds of prey species, the long-winged harrier possesses sharp talons used to catch their prey on the ground or in the air, and their legs are bright orange. The species also shares the large hooked beak of other birds of prey, which it uses to tear apart meat once it catches its prey.
The long-winged harrier is part of the only diurnal bird of prey group to possess a facial disc
In ornithology, the facial disc is the concave collection of feathers on the face of some birds—most notably owls—surrounding the eyes. The concavity of the facial disc forms a circular paraboloid that collects sound waves and directs those ...
. The facial disc is formed by a group of easily recognizable group of feathers that form a disc shape around the bird's face. These feathers can be raised in response to sounds and can improve the bird's hearing when it is hunting by triangulating sounds in its ear holes.
Little is known of the long-winged harrier's vocalization, as is not a very vocal bird. It will call mostly when in flight and some variation of the bird's call include a fast succession of "klee-klee-klee", and a more quiet "thrill" with a quickly descending pitch.
Distribution and habitat
The long-winged harrier's year-round range encompasses most of eastern South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
, including eastern Argentina
Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, Uruguay
Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
, Brazil
Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
and Paraguay
Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
, and extends up to Colombia, Venezuela
Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
, and the northern tips of Guyana, Suriname and French Guiana
French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas department/region and single territorial collectivity of France on the northern Atlantic coast of South America in the Guianas. ...
. The species has also been observed occasionally in extant areas such as Bolivia, Peru
, image_flag = Flag of Peru.svg
, image_coat = Escudo nacional del Perú.svg
, other_symbol = Great Seal of the State
, other_symbol_type = National seal
, national_motto = "Firm and Happy f ...
, Chile
Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
, Trinidad and Tobago
Trinidad and Tobago (, ), officially the Republic of Trinidad and Tobago, is the southernmost island country in the Caribbean. Consisting of the main islands Trinidad and Tobago, and numerous much smaller islands, it is situated south of ...
, and all the way down to Tierra del Fuego
Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla ...
in southern Argentina
Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. It is vagrant to Panama
Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Co ...
and the Falkland Islands
The Falkland Islands (; es, Islas Malvinas, link=no ) is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and about from Cape Dubouze ...
.
This species occupies large grasslands and agricultural fields, where their wide-range active hunting technique is very effective. Their habitat also includes savanna
A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
, marsh
A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found a ...
es and wetland
A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The p ...
s of South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
.
Behavior and ecology
Food and feeding
As all of the extended members of the family Accipitridae
The Accipitridae is one of the three families within the order Accipitriformes, and is a family of small to large birds with strongly hooked bills and variable morphology based on diet. They feed on a range of prey items from insects to medium-s ...
, the long-winged harrier is carnivorous, feeding on a large variety of animals due to its widespread distribution. Their diet includes small mammals (such as cavies
Caviidae, the cavy family, is composed of rodents native to South America and includes the domestic guinea pig, wild cavies, and the largest living rodent, the capybara. They are found across South America in open areas from moist savanna to tho ...
), amphibians (such as frog
A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-frog" ''Triadobatrachus'' is ...
s), reptiles (such as lizards) and birds (such as crakes and young coot
Coots are medium-sized water birds that are members of the rail family, Rallidae. They constitute the genus ''Fulica'', the name being the Latin term for "coot". Coots have predominantly black plumage, and—unlike many rails—they are usually ...
s), with birds consisting the majority of their food intake.[Ferguson-Lees, J. & Christie, D.A. & Franklin, K. & Mead, D. & Burton, P.. (2001). Raptors of the world. Helm Identification Guides.] It will also feed on bird eggs, varying from taking the whole ball-nests of water-tyrants to raiding heronries
A heronry, sometimes called a heron rookery, is a breeding ground for herons.
Notable heronries
Although their breeding territories are often on more protected small islands in lakes or retention ponds, herons breed in heronries (or also calle ...
. Its hunting strategy differs vastly from other hawks sharing the same habitat such as the roadside hawk
The roadside hawk (''Rupornis magnirostris'') is a relatively small bird of prey found in the Americas. This vocal species is often the most common raptor in its range. It has many subspecies and is now usually placed in the monotypic genus ''Ru ...
(''Rupornis magnirostris''). Instead of opting for a passive hunting strategy consisting of the use of high perches to locate their prey, the long-winged harrier will fly slowly in large circles above its range in order to find its prey, using its keen vision and hearing.
Breeding
The long-winged harrier nests on the ground, building its nest between September and October in the grasslands of its habitat. The nests are usually made out of rushes and grass, and are built no more than 3 meters up the ground and no less than 1 meter away from bodies of water. Their breeding season can vary vastly across their range, but it will usually take place in the summer months (September to January). During their reproduction cycle, the nests will usually hold between 3 and 4 eggs per clutch.
Conservation
Although the species population has been found to be declining, the extended range of the long-winged harrier allows it to be classified as a species of 'least concern' by the IUCN Red List
The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
. Their population decline has been attributed to the degradation of its habitat, most notably wetlands, due to drainage, pollution, and other human-caused circumstances. No conservation efforts are currently in action concerning the long-winged harrier as an individual species, but their range occurs in a wide variety of conservation sites and protected areas.
References
External links
eBird
Range and other media, including vocalization
Peruaves.org
Long-winged harrier (''Circus buffoni'')
{{Taxonbar, from=Q520910
long-winged harrier
Birds of Colombia
Birds of Venezuela
Birds of Trinidad and Tobago
Birds of the Guianas
Birds of Brazil
Birds of Paraguay
Birds of Argentina
Birds of prey of South America
long-winged harrier
long-winged harrier
Taxonomy articles created by Polbot