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The long-billed hermit (''Phaethornis longirostris'') is a
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in the family
Trochilidae Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics arou ...
, the
hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...
s. It is found from central
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
south through
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
,
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
and
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
into
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
.


Taxonomy and systematics

It has often been considered
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organ ...
with what is now the
long-tailed hermit The long-tailed hermit (''Phaethornis superciliosus'') is a large hummingbird that is a resident breeder in Venezuela, the Guianas, and north-eastern Brazil. This species was formerly referred to as the eastern long-tailed hermit. Taxonomy The lo ...
, ''P. superciliosus'', which is found east of the
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
. The two populations were called the western and eastern long-tailed hermits; the "eastern" was dropped after the split into two species and the renaming of the western population.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 August 2021. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved August 24, 2021 The South American Classification Committee (SACC) of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its m ...
, the
International Ornithological Committee The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...
(IOC), and the
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 2022 ...
assign these four subspecies to the long-billed hermit:Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021 *''P. l. longirostris'' Delattre (1843) *''P. l. cephalus'' Bourcier & Mulsant (1848) *''P. l. susurrus''
Bangs Bang or bangs may refer to: Products * M1922 Bang rifle, a US semi-automatic rifle designed by Søren Hansen Bang * Bang, a List of model car brands, model car brand * Bang (beverage), an energy drink Geography * Bang, Lorestan, a village in I ...
(1901)
*''P. l. baroni''
Hartert Ernst Johann Otto Hartert (29 October 1859 – 11 November 1933) was a widely published German ornithologist. Life and career Hartert was born in Hamburg, Germany on 29 October 1859. In July 1891, he married the illustrator Claudia Bernadine E ...
(1897)
However,
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
's ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. T ...
'' (HBW) treats ''P. l. baroni'' as a separate species, "Ecuadorian hermit".HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved May 27, 2021 The SACC in 2006 had rejected that treatment. Other subspecies within ''longirostrus'' and ''cephalus'' have been proposed but have not been accepted.del Hoyo, J., C. Hinkelmann, N. Collar, P. F. D. Boesman, and G. M. Kirwan (2020). Long-billed Hermit (''Phaethornis longirostris''), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.lobher.01 retrieved December 9, 2021


Description

The long-billed hermit is long. Males weigh and females . The nominate subspecies ''P. l. longirostris'' has greenish-brown upperparts and brownish to grayish underparts. Its uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are sm ...
have dark and light ochre bands and the central tail feathers are long with long white tips. It has a long decurved bill, with the female's being shorter but more curved than the male's. ''P. l. cephalus'' is similar but its underparts become more ochraceous towards the southern part of its range. ''P. l. susurrus'' is larger than ''cephalus'' but similarly colored. ''P. l. baroni''s upperparts are dull metallic green and its belly is whiter than those of the other subspecies.


Distribution and habitat

The subspecies of long-billed hermit are found thus: *''P. l. longirostris'', from Mexico's
Oaxaca Oaxaca ( , also , , from nci, Huāxyacac ), officially the Free and Sovereign State of Oaxaca ( es, Estado Libre y Soberano de Oaxaca), is one of the 32 states that compose the political divisions of Mexico, Federative Entities of Mexico. It is ...
and
Chiapas Chiapas (; Tzotzil language, Tzotzil and Tzeltal language, Tzeltal: ''Chyapas'' ), officially the Free and Sovereign State of Chiapas ( es, Estado Libre y Soberano de Chiapas), is one of the states that make up the Political divisions of Mexico, ...
states south through
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
into northern
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. The republic of Honduras is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Oce ...
*''P. l. cephalus'', from eastern Honduras through
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the cou ...
,
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
, and
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
into northwestern Colombia as far south as Santander Department *''P. l. susurrus'', the isolated
Sierra Nevada de Santa Marta The Sierra Nevada de Santa Marta (English: ''Snow-Covered Mountain Range of Saint Martha'') is an isolated mountain range in northern Colombia, separate from the Andes range that runs through the north of the country. Reaching an elevation of ...
in northeastern Colombia *''P. l. baroni'', from Ecuador's western
Esmeraldas Province Esmeraldas () is a province in northwestern Ecuador. The capital is Esmeraldas. The province is home to the Afro-Ecuadorian culture. Demographics Ethnic groups as of the Ecuadorian census of 2010: *Mestizo 44.7% * Afro-Ecuadorian 43.9% *W ...
south into Peru's Tumbes and
Piura Piura is a city in northwestern Peru located in the Sechura Desert on the Piura River. It is the capital of the Piura Region and the Piura Province. Its population was 484,475 as of 2017. It was here that Spanish Conqueror Francisco Pizarro fou ...
departments. The species inhabits the understory and edges of a variety of landscapes including
rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...
, tall
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has re-grown after a timber harvest or clearing for agriculture, until a long enough period has passed so that the effects of the disturbance are no longer evident. ...
, humid
semi-deciduous Semi-deciduous or semi-evergreen is a botanical term which refers to plants that lose their foliage for a very short period, when old leaves fall off and new foliage growth is starting. This phenomenon occurs in tropical and sub-tropical woody spec ...
forest,
cloudforest A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest (TMCF), is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud c ...
, and
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
. It has been recorded from sea level to in Costa Rica, to in eastern Mexico, to in northern Colombia, to in Peru, and to at least and possibly to in Ecuador.


Behavior


Movement

The long-billed hermit is not known to make large-scale movements but is thought to wander short distances.


Feeding

The long-billed hermit is a " trap-line" feeder like other hermit hummingbirds, visiting a circuit of a variety of flowering plants for nectar. Examples include ''
Heliconia ''Heliconia'', derived from the Greek word (), is a genus of flowering plants in the monotypic family Heliconiaceae. Most of the ca 194 known species are native to the tropical Americas, but a few are indigenous to certain islands of the we ...
'', ''
Costus ''Costus'' is a group of herbaceous perennial plants in the family Costaceae, described by Linnaeus as a genus in 1753. It was formerly known as ''Hellenia'' after the Finnish botanist Carl Niclas von Hellens. It is widespread through tropica ...
'', '' Aphelandra'', and '' Pasiflora''. It also consumes small
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s.


Breeding

The long-billed hermit's breeding seasons vary considerably throughout its range, from April to July in Mexico, May to September in Panama, and January to April or May in Colombia. The nest is a cone-shaped cup made of plant fibers and spider silk suspended from the underside of a drooping leaf. The clutch size is two eggs; The incubation period is 17 to 18 days with
fledging Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
22 to 23 days after hatch. During the breeding season, male long-billed hermits sing in communal
lek Lek or LEK may refer to: * Lek mating, mating in a lek, a type of animal territory in which males of a species gather * Albanian lek, the currency of Albania * Lek (magazine), a Norwegian softcore pornographic magazine * Lek (pharmaceutical comp ...
s of up to 25 birds, and also wiggle their long tails in display. Competitive lek singing can occupy half of the daylight hours to attract females. In addition, mature males have a longer
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
than females, and appear to use its
dagger A dagger is a fighting knife with a very sharp point and usually two sharp edges, typically designed or capable of being used as a thrusting or stabbing weapon.State v. Martin, 633 S.W.2d 80 (Mo. 1982): This is the dictionary or popular-use de ...
-like tip as a secondary sexual trait to defend against other males at the lek.


Vocalization

The songs of the three northern subspecies of long-billed hermit are "a continuous series of single, piercing, usually upslurred 'sweeup' notes". Their call is "a sharp, explosive 'week!'" that is often given in flight. The song of ''P. l. baroni'' is "a continuous series of single, sparrow-like, chipping notes...'tchee..tchee..tchee..'." Its call is "a thin 'seep'" also given in flight.


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
follows HBW taxonomy, and so has separately assessed the long-billed hermit and "Ecuadorian hermit" as being of Least Concern. The population sizes and trends of both are unknown. ''P. l. susurrus'' has a small range that potentially places it at risk. ''P. l. cephalus'' occurs in many protected areas in Costa Rica. ''P. l. baroni'' is considered fairly common throughout its range and occurs in some protected areas.


References


External links


Article
AOCR


Further reading

* Hilty, '' Birds of Venezuela'', * Stiles and Skutch, ''A guide to the birds of Costa Rica'' {{Taxonbar, from=Q1055821
long-billed hermit The long-billed hermit (''Phaethornis longirostris'') is a bird in the family Trochilidae, the hummingbirds. It is found from central Mexico south through Central America, Colombia and Ecuador into Peru. Taxonomy and systematics It has oft ...
Hummingbird species of Central America Birds of Mexico Birds of Belize Birds of Guatemala Birds of Honduras Birds of Nicaragua Birds of Costa Rica Birds of Panama Birds of Colombia Hummingbird species of South America long-billed hermit'