Locustellidae is a newly recognized

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

of small insectivorous

songbird A songbird is a bird belonging to the suborder Passeri of the perching birds (Passeriformes). Another name that is sometimes seen as the scientific or vernacular name is Oscines, from Latin ''oscen'', "songbird". The Passeriformes contains 500 ...

s ("

warblers Various Passeriformes (perching birds) are commonly referred to as warblers. They are not necessarily closely related to one another, but share some characteristics, such as being fairly small, vocal, and insectivorous. Sylvioid warblers Th ...

"), formerly placed in the

Old World warbler Old World warblers are a large group of birds formerly grouped together in the bird family Sylviidae. The family held over 400 species in over 70 genera, and were the source of much taxonomic confusion. Two families were split out initially, the ci ...

wastebin A waste container, also known as a dustbin, garbage can, and trash can is a type of container that is usually made out of metal or plastic. The words "rubbish", "basket" and "bin" are more common in British English usage; "trash" and "can" a ...

. It contains the

grass warbler The grass warblers are small passerine birds belonging to the genus ''Locustella''. Formerly placed in the paraphyletic "Old World warbler" assemblage, they are now considered the northernmost representatives of a largely Gondwanan family, the L ...

s, grassbirds, and the ''

Bradypterus ''Bradypterus'' is a genus of small insectivorous songbirds ("warblers") in the newly recognized grass warbler family (Locustellidae). They were formerly placed in the Sylviidae, which at that time was a wastebin taxon for the warbler-like Syl ...

'' "

bush warbler Bush-warblers (or bush warblers) are small insectivorous songbirds of the genera ''Cettia'', ''Horornis'', and ''Bradypterus''. They were formerly placed in the "wastebin" Old World warbler family. None of the genera as traditionally delimited ...

s". These birds occur mainly in

Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

, and the

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

n region. The family name is sometimes given as Megaluridae, but Locustellidae has priority. The

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

are smallish birds with tails that are usually long and pointed; the

scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

of the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

''Megalurus'' in fact means "the large-tailed one" in plain English. They are less wren-like than the typical shrub-warblers (''

Cettia ''Cettia'' is a genus of small insectivorous songbirds (" warblers") which make up the core of the newly recognized family Cettiidae. They were formerly placed in the Sylviidae, which at that time was a wastebin taxon for the warbler-like Sylv ...

''), but they are similarly drab brownish or buffy all over. They tend to be larger and slimmer than ''Cettia'' though, and many have bold dark streaks on wings and/or underside. Most live in scrubland and frequently hunt food by clambering through thick tangled growth or pursuing it on the ground; they are perhaps the most

terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...

of the "warblers". Very unusual for

Passeriformes A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...

, the beginning of an

evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...

towards flightlessness is seen in some

taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

. Among the "warbler and babbler"

superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...

Sylvioidea Sylvioidea is a superfamily of passerine birds, one of at least three major clades within the Passerida along with the Muscicapoidea and Passeroidea. It contains about 1300 species including the Old World warblers, Old World babblers, swallows ...

, the Locustellidae are closest to the

Malagasy warblers The Tetrakas and allies are a newly validated family of songbirds. They were formally named Bernieridae in 2010. The family currently consists of eleven species (in eight genera) of small forest birds. These birds are all endemic to Madagascar ...

, another newly recognized (and hitherto unnamed) family. The

black-capped donacobius The black-capped donacobius (''Donacobius atricapilla'') is a conspicuous, vocal South American bird. It is found in tropical swamps and wetlands in Argentina, Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Paraguay, Peru, Suriname, ...

(''Donacobius atricapillus'') is a

South American South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...

relative derived from the same ancestral stock and not a wren as was long believed. A comprehensive

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

study of the grassbird family Locustellidae published in 2018 found that many of the genera, as then defined, were non-

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...

. The resulting revision of the genus level taxonomy involved many changes including the resurrection of the genera '' Poodytes'' and '' Cincloramphus'' as well as the erection of a new genus ''

Helopsaltes ''Helopsaltes'' is a genus of passerine birds in the grassbird family Locustellidae. A comprehensive molecular phylogenetic study of the grassbird family Locustellidae published in 2018 found that the genus '' Locustella'' consisted of two disti ...

''. The former genera ''Megalurulus'' and ''Buettikoferella'' become

junior synonyms The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linn ...

of ''Cincloramphus''.

Genera

The family contains 67 species divided into 11 genera. * '' Robsonius'' – ground warblers (3 species) * ''

'' – grasshopper warblers (6 species) * '' Locustella'' – grass warblers (23 species) * ''

'' – megalurid bush warblers (12 species) * '' Elaphrornis'' – Sri Lanka bush warbler * '' Schoenicola'' – wide-tailed grassbirds (2 species) * '' Poodytes'' – grassbirds (5 species including one extinct) * '' Malia'' – malia * '' Cincloramphus'' – (12 species) * '' Megalurus'' – striated grassbird * '' Catriscus'' – fan-tailed grassbird The relationships between the genera is shown in the following cladogram. It is based on a 2018 study by

Per Alström ''Per'' Johan Alström (born 9 April 1961) is a Swedish Professor of ornithology. He researches in taxonomy, systematics, and evolution, with birds in Asia as a specialty. Alström works at the Department of Ecology and Genetics (Animal Ecology) a ...

and coworkers.

References

{{Taxonbar, from=Q774034 Bird families Taxa named by Charles Lucien Bonaparte