Llewellya Hillis
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Llewellya Williams Hillis (1930 – March 23, 2019), later Llewellya Hillis-Colinvaux, was a Canadian-born American marine biologist.


Early life

Llewellya Hillis was born in
Windsor, Ontario Windsor is a city in southwestern Ontario, Canada, on the south bank of the Detroit River directly across from Detroit, Michigan, United States. Geographically located within but administratively independent of Essex County, it is the souther ...
and raised in Walkerville. She graduated from
Walkerville Collegiate Institute Walkerville Collegiate Institute (WCI) is a secondary school (grades 9 to 12) in the Walkerville area of Windsor, Ontario and managed by the Greater Essex County District School Board. It is the home of the Walkerville Centre for the Creative A ...
. Her father Llewellyn Hillis worked at an automotive plant, and her mother Pearl Evelina Hillis was a teacher. She earned her bachelor's degree at Queen's University in 1952. In 1957 she completed her doctoral work in botany at the
University of Michigan , mottoeng = "Arts, Knowledge, Truth" , former_names = Catholepistemiad, or University of Michigania (1817–1821) , budget = $10.3 billion (2021) , endowment = $17 billion (2021)As o ...
; her thesis titled "A Revision of the Genus Halimeda (order Siphonales)" was published in 1959. As a graduate student, she did research at the
Marine Biological Laboratory The Marine Biological Laboratory (MBL) is an international center for research and education in biological and environmental science. Founded in Woods Hole, Massachusetts, in 1888, the MBL is a private, nonprofit institution that was independent ...
at
Woods Hole, Massachusetts Woods Hole is a census-designated place in the town of Falmouth in Barnstable County, Massachusetts, United States. It lies at the extreme southwest corner of Cape Cod, near Martha's Vineyard and the Elizabeth Islands. The population was 781 at ...
.


Career

Hillis held a post-doctoral appointment at the
University of New Brunswick The University of New Brunswick (UNB) is a public university with two primary campuses in Fredericton and Saint John, New Brunswick. It is the oldest English-language university in Canada, and among the oldest public universities in North Americ ...
, before joining the botany faculty at
Ohio State University The Ohio State University, commonly called Ohio State or OSU, is a public land-grant research university in Columbus, Ohio. A member of the University System of Ohio, it has been ranked by major institutional rankings among the best publ ...
in 1964 (she transferred to the zoology faculty in 1972). "It was a ten-year fight to be recognized. No recognition has come to me as easily as it has to my male counterparts," she said of her academic career. "However, the progress that has been made is now so firmly entrenched that it will be hard to dismantle it completely." Though landlocked Ohio was not the ideal place to be a marine biologist, she continued her work on coral reef algae, especially in the genus ''
Halimeda ''Halimeda'' is a genus of green macroalgae. The algal body (thallus) is composed of calcified green segments. Calcium carbonate is deposited in its tissues, making it inedible to most herbivores. However one species, '' Halimeda tuna'', was desc ...
''. She secured funding from the U.S.
Office of Naval Research The Office of Naval Research (ONR) is an organization within the United States Department of the Navy responsible for the science and technology programs of the U.S. Navy and Marine Corps. Established by Congress in 1946, its mission is to plan ...
and from the
National Science Foundation The National Science Foundation (NSF) is an independent agency of the United States government that supports fundamental research and education in all the non-medical fields of science and engineering. Its medical counterpart is the National I ...
. She imported seawater to Ohio to cultivate a seaweed colony for study. In 1976, she traveled to
Enewetak Atoll Enewetak Atoll (; also spelled Eniwetok Atoll or sometimes Eniewetok; mh, Ānewetak, , or , ; known to the Japanese as Brown Atoll or Brown Island; ja, ブラウン環礁) is a large coral atoll of 40 islands in the Pacific Ocean and with it ...
to find ''Halimeda'' in a nuclear bomb crater. Hillis and Colinvaux left Ohio in the 1990s and continued their research at the
Smithsonian Tropical Research Institute The Smithsonian Tropical Research Institute (STRI, es, Instituto Smithsonian de Investigaciones Tropicales) is located in Panama and is the only bureau of the Smithsonian Institution based outside of the United States. It is dedicated to understa ...
in Panama. She published a monograph on ''Halimeda'' in 1980. Hillis held fellowships at the
British Museum The British Museum is a public museum dedicated to human history, art and culture located in the Bloomsbury area of London. Its permanent collection of eight million works is among the largest and most comprehensive in existence. It docum ...
(1971) and the
Bunting Institute The Radcliffe Institute for Advanced Study at Harvard University—also known as the Harvard Radcliffe Institute—is a part of Harvard University that fosters interdisciplinary research across the humanities, sciences, social sciences, arts, a ...
(1985-1987). Two coral reef species are named for her: ''Carpathea llewellyae'' and ''Leckhamptonella llewellyae''.


Selected Publications

* Ecology and Taxonomy of Halimeda: Primary Producer of Coral Reefs. Llewellya Hillis-Colinvaux. Advances in Marine Biology. Volume 17, 1980, Pages 1-327. * Geology and biological zonation of the reef slope, 50–360 m depth at Enewetak Atoll, Marshall Islands. Colin, Patrick L.; Devaney, Dennis M.; Hillis-Colinvaux, Llewellya; Suchanek, Thomas H.; and Harrison, John T.
Bulletin of Marine Science The ''Bulletin of Marine Science'' is a peer-reviewed scientific journal published by the Rosenstiel School of Marine and Atmospheric Science of the University of Miami. The journal was established in 1951 as the ''Bulletin of Marine Science of ...
, Volume 38, Number 1, January 1986, pp. 111-128(18). * Electron microscope study of calcification in the alga Halimeda (order Siphonales). Karl M. Wilbur, Llewellya Hillis Colinvaux and Norimitsu Watabe.
Phycologia The International Phycological Society is a learned society of phycologists. It was established in 1960. The Society publishes a bimonthly academic journal An academic journal or scholarly journal is a periodical publication in which scholarsh ...
Volume 8, 1969 - Issue 1. * Halimeda growth and diversity on the deep fore-reef of Enewetak Atoll. Llewellya Hillis Colinvaux.
Coral Reefs A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
. * Deep water populations of Halimeda in the economy of an atoll. Hillis-Colinvaux, Llewellya.
Bulletin of Marine Science The ''Bulletin of Marine Science'' is a peer-reviewed scientific journal published by the Rosenstiel School of Marine and Atmospheric Science of the University of Miami. The journal was established in 1951 as the ''Bulletin of Marine Science of ...
, Volume 38, Number 1, January 1986, pp. 155-169(15).


Personal life

Hillis married fellow biologist Paul Colinvaux in 1961. They had two children, Roger and Catherine. They retired to
Cape Cod Cape Cod is a peninsula extending into the Atlantic Ocean from the southeastern corner of mainland Massachusetts, in the northeastern United States. Its historic, maritime character and ample beaches attract heavy tourism during the summer mont ...
, and were active in the Woods Hole community in their later years. Hillis was widowed when Colinvaux died in 2016, and she died in 2019, on Cape Cod, aged 89 years.


References

* {{DEFAULTSORT:Hillis, Llewellya 1930 births 2019 deaths American women scientists American phycologists Women phycologists American marine biologists Canadian women scientists Canadian marine biologists Ohio State University faculty University of Michigan alumni People from Windsor, Ontario Scientists from Ontario American women academics 21st-century American women