The little penguin (''Eudyptula minor'') is a species of
penguin
Penguins (order (biology), order List of Sphenisciformes by population, Sphenisciformes , family (biology), family Spheniscidae ) are a group of Water bird, aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: on ...
from
New Zealand
New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
. They are commonly known as little blue penguins or blue penguins owing to their
slate
Slate is a fine-grained, foliated, homogeneous metamorphic rock derived from an original shale-type sedimentary rock composed of clay or volcanic ash through low-grade regional metamorphism. It is the finest grained foliated metamorphic rock. ...
-blue plumage and are also known by their
Māori
Māori or Maori can refer to:
Relating to the Māori people
* Māori people of New Zealand, or members of that group
* Māori language, the language of the Māori people of New Zealand
* Māori culture
* Cook Islanders, the Māori people of the C ...
name .
The
Australian little penguin
The Australian little penguin (''Eudyptula novaehollandiae''), also called the fairy penguin, is a species of penguin from Australia and the Otago region of New Zealand. The species was described as ''Spheniscus'' ''novaehollandiae'' in 1826. ...
(''Eudyptula novaehollandiae'') from
Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
and the
Otago
Otago (, ; mi, Ōtākou ) is a region of New Zealand located in the southern half of the South Island administered by the Otago Regional Council. It has an area of approximately , making it the country's second largest local government reg ...
region of New Zealand is considered a separate species by a 2016 study
and a 2019 study.
Taxonomy
The little penguin was first described by German
naturalist Johann Reinhold Forster
Johann Reinhold Forster (22 October 1729 – 9 December 1798) was a German Reformed (Calvinist) pastor and naturalist of partially Scottish descent who made contributions to the early ornithology of Europe and North America. He is best known ...
in 1781. Several
subspecies
In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are known, but a precise classification of these is still a matter of dispute. The
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
s of the subspecies ''E. m. variabilis'' and ''Eudyptula minor chathamensis'' are in the collection of the
Museum of New Zealand Te Papa Tongarewa. The
white-flippered penguin (''E. m. albosignata'' or ''E. m. minor
morpha albosignata'') is currently considered by most
taxonomists
In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ...
to be a colour
morph
Morph may refer to:
Biology
* Morph (zoology), a visual or behavioral difference between organisms of distinct populations in a species
* Muller's morphs, a classification scheme for genetic mutations
* "-morph", a suffix commonly used in tax ...
or subspecies of ''Eudyptula minor.'' In 2008,
Shirihai treated the little penguin and white-flippered penguin as
allospecies. However, as of 2012, the
IUCN and
BirdLife International
BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
consider the white-flippered penguin to be a subspecies or morph of the little penguin.
Little penguins from New Zealand and Australia were once considered to be the same species, called ''Eudyptula minor''. Analysis of
mtDNA
Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
in 2002 revealed two
clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
s in ''
Eudyptula
The genus ''Eudyptula'' ("good little diver") contains two species of penguin, found in southern Australia, Tasmania, and New Zealand (including the Chatham Islands). They are commonly known as the little penguin, little blue penguin, or, in Aus ...
'': one containing little penguins of New Zealand's
North Island
The North Island, also officially named Te Ika-a-Māui, is one of the two main islands of New Zealand, separated from the larger but much less populous South Island by the Cook Strait. The island's area is , making it the world's 14th-largest ...
,
Cook Strait
Cook Strait ( mi, Te Moana-o-Raukawa) separates the North and South Islands of New Zealand. The strait connects the Tasman Sea on the northwest with the South Pacific Ocean on the southeast. It is wide at its narrowest point,McLintock, A ...
and
Chatham Island
Chatham Island ( ) (Moriori: ''Rēkohu'', 'Misty Sun'; mi, Wharekauri) is by far the largest island of the Chatham Islands group, in the south Pacific Ocean off the eastern coast of New Zealand's South Island. It is said to be "halfway bet ...
, as well as the white-flippered penguin, and a second containing little penguins of
Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
and the
Otago
Otago (, ; mi, Ōtākou ) is a region of New Zealand located in the southern half of the South Island administered by the Otago Regional Council. It has an area of approximately , making it the country's second largest local government reg ...
region of New Zealand.
[Banks, Jonathan C.; Mitchell, Anthony D.; Waas, Joseph R. & Paterson, Adrian M. (2002): An unexpected pattern of molecular divergence within the blue penguin (''Eudyptula minor'') complex. ''Notornis'' 49(1): 29–38]
PDF fulltext
/ref> Preliminary analysis of braying calls and cluster analysis
Cluster analysis or clustering is the task of grouping a set of objects in such a way that objects in the same group (called a cluster) are more similar (in some sense) to each other than to those in other groups (clusters). It is a main task of ...
of morphometrics
Morphometrics (from Greek μορϕή ''morphe'', "shape, form", and -μετρία ''metria'', "measurement") or morphometry refers to the quantitative analysis of ''form'', a concept that encompasses size and shape. Morphometric analyses are co ...
partially supported these results.[ A 2016 study described the Australian little penguin as a new and separate species, '']Eudyptula novaehollandiae
The Australian little penguin (''Eudyptula novaehollandiae''), also called the fairy penguin, is a species of penguin from Australia and the Otago region of New Zealand. The species was described as ''Spheniscus'' ''novaehollandiae'' in 1826. It ...
''. ''E. minor'' is endemic to New Zealand, while ''E. novaehollandiae'' is found in Australia and Otago. A 2019 study supported the recognition of ''E. minor'' and ''E. novaehollandiae'' as separate species.
Description
Like those of all penguins
Penguins (order Sphenisciformes , family Spheniscidae ) are a group of aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: only one species, the Galápagos penguin, is found north of the Equator. Highly adapt ...
, the wings of ''Eudyptula
The genus ''Eudyptula'' ("good little diver") contains two species of penguin, found in southern Australia, Tasmania, and New Zealand (including the Chatham Islands). They are commonly known as the little penguin, little blue penguin, or, in Aus ...
'' species have developed into flippers used for swimming.
''Eudyptula'' species typically grow to between tall and on average weigh 1.5 kg (3.3 lb). The head and upper parts are blue in colour, with slate-grey ear coverts fading to white underneath, from the chin to the belly. Their flippers are blue in colour. The dark grey-black beak is 3–4 cm long, the irises pale silvery- or bluish-grey or hazel, and the feet pink above with black soles and webbing. An immature individual will have a shorter bill and lighter upperparts.
Like most seabirds, the ''Eudyptula'' species have a long lifespan. The average for the species is 6.5 years, but flipper ringing experiments show that in very exceptional cases they may live up to 25 years in captivity.
''Eudyptula minor'' does not have the distinct bright blue feathers that distinguish ''Eudyptula novaehollandiae.'' In addition, the vocalisation patterns of the New Zealand lineage located on Tiritiri Matangi Island
Tiritiri Matangi Island is located in the Hauraki Gulf of New Zealand, east of the Whangaparaoa Peninsula in the North Island and north east of Auckland. The island is an open nature reserve managed by the Supporters of Tiritiri Matangi Incorp ...
vary from the Australian lineage located in Oamaru
Oamaru (; mi, Te Oha-a-Maru) is the largest town in North Otago, in the South Island of New Zealand, it is the main town in the Waitaki District. It is south of Timaru and north of Dunedin on the Pacific coast; State Highway 1 and the railway ...
. Females are known to prefer the local call of the New Zealand lingeage.
There are also behavioural differences that help differentiate these penguins. Those of the Australian lineage will swim together in a large group after dusk and walk along the shore to reach their nesting sites. This may be an effective predator avoidance strategy by traveling in a large group simultaneously. This has not been seen by those of the New Zealand lineage. ''Eudyptula'' ''minor'' only recently encountered terrestrial vertebrate predators, while ''Eudyptula novaehollandiae'' would have had to deal with carnivorous marsupials.
Also, ''Eudyptula novaehollandiae'' located in Australia will double brood. Birds will double brood by laying another clutch of eggs in hopes to increase their reproductive success. They complete this after the first clutch has successfully fledged. They may also do this due to the increasing sea surface temperatures and changing sources of food that are available. This behaviour has never been observed by those in New Zealand.
Distribution and habitat
New Zealand
''Eudyptula minor'' breeds along most of the coastline of New Zealand
New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, including the Chatham Islands
The Chatham Islands ( ) (Moriori: ''Rēkohu'', 'Misty Sun'; mi, Wharekauri) are an archipelago in the Pacific Ocean about east of New Zealand's South Island. They are administered as part of New Zealand. The archipelago consists of about te ...
. However, ''Eudyptula minor'' does not occur in Otago
Otago (, ; mi, Ōtākou ) is a region of New Zealand located in the southern half of the South Island administered by the Otago Regional Council. It has an area of approximately , making it the country's second largest local government reg ...
, which is located on the east coast of New Zealand's South Island
The South Island, also officially named , is the larger of the two major islands of New Zealand in surface area, the other being the smaller but more populous North Island. It is bordered to the north by Cook Strait, to the west by the Tasman ...
. The Australian species ''Eudyptula novaehollandiae
The Australian little penguin (''Eudyptula novaehollandiae''), also called the fairy penguin, is a species of penguin from Australia and the Otago region of New Zealand. The species was described as ''Spheniscus'' ''novaehollandiae'' in 1826. It ...
'' occurs in Otago. ''E. novaehollandiae'' was originally endemic to Australia. Using ancient-DNA analysis and radiocarbon dating using historical, pre-human, as well as archaeological ''Eudyptula'' remains, the arrival of the Australian species in New Zealand was determined to have occurred roughly between AD 1500 and 1900. When the ''E. minor'' population declined in New Zealand, it left a genetic opening for ''E. novaehollandiae''. The decrease of ''E. minor'' was most likely due to anthropogenic effects, such as being hunted by humans as well as introduced predators, including dogs brought from overseas.
It has been determined that the population of ''Eudyptula novaehollandiae'' in Otago arrived even more recently than previously estimated due to mulitlocus coalescent analyses.
Overall, little penguin populations in New Zealand have been decreasing. Some colonies have become extinct, and others continue to be at risk. Some new colonies have been established in urban areas. The species is not considered endangered in New Zealand, with the exception of the white-flippered subspecies found only on Banks Peninsula
Banks Peninsula is a peninsula of volcanic origin on the east coast of the South Island of New Zealand. It has an area of approximately and encompasses two large harbours and many smaller bays and coves. The South Island's largest cit ...
and nearby Motunau Island
Motunau Island is a small, , island nature reserve lying off the coast of New Zealand's South Island, at the northern end of Pegasus Bay, south of the mouth of the Motunau River. The reserve is managed by the Department of Conservation (New Ze ...
. Since the 1960s, the mainland population has declined by 60-70%; though a small increase has occurred on Motunau Island. A colony exists in Wellington
Wellington ( mi, Te Whanganui-a-Tara or ) is the capital city of New Zealand. It is located at the south-western tip of the North Island, between Cook Strait and the Remutaka Range. Wellington is the second-largest city in New Zealand by me ...
Harbor on Matiu / Somes Island
Matiu / Somes Island is the largest of three islands in the northern half of Wellington Harbour, New Zealand. It is the former site of military and quarantine internments, as well as animal quarantine until 1995. Since 1995 it has been designat ...
.
Outside of Australasia
''Eudyptula'' species have also been reported from Chile
Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
, where they are known as ''pingüino pequeño'' or ''pingüino azul.'' Sightings include Isla Chañaral
Isla or ISLA may refer to:
Organizations
* International Securities Lending Association, a trade association
* International School of Los Angeles
* International Bilingual School, later named International School of Los Angeles
People
* Isla (g ...
1996 and Playa de Santo Domingo, San Antonio
("Cradle of Freedom")
, image_map =
, mapsize = 220px
, map_caption = Interactive map of San Antonio
, subdivision_type = Country
, subdivision_name = United States
, subdivision_type1= U.S. state, State
, subdivision_name1 = Texas
, s ...
, 16 March 1997. ''Eudyptula'' species have also been reported from South Africa
South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
, It is unclear whether these birds were vagrants
Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, tempora ...
.
Behaviour
Feeding
Little penguins feed by hunting small clupeoid fish, cephalopod
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head ...
s, and crustacean
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s, for which they travel and dive quite extensively including to the sea floor. Important little penguin prey items include arrow squid, slender sprat, Graham's gudgeon, red cod
''Pseudophycis'' is a genus of codlings of the family Moridae found around New Zealand and Southern Australia.
Species
The currently recognized species in this genus are:
* '' Pseudophycis bachus'' ( J. R. Forster, 1801) (red codling)
* ''Pseud ...
, and ahuru
The ahuru (''Auchenoceros punctatus'') is a species of morid cod found in waters off the eastern coast of New Zealand and to the south as well as in the Cook Strait. It is found at depths from . This species grows to in total length. It is t ...
.[Flemming, S.A. (2013)]
. ''In Miskelly, C.M. (ed.)'' ''New Zealand Birds Online''
Threats
Protestors have opposed the development of a marina at Kennedy Point, Waiheke Island
Waiheke Island (; Māori: ) is the second-largest island (after Great Barrier Island) in the Hauraki Gulf of New Zealand. Its ferry terminal in Matiatia Bay at the western end is from the central-city terminal in Auckland.
It is the most pop ...
in New Zealand for the risk it poses to little penguins and their habitat. Protesters claimed that they exhausted all legal means to oppose the project and have had to resort to occupation and non-violent resistance. Several arrests have been made for trespassing.
Introduced predators
Introduced mammalian predators present the greatest terrestrial risk to little penguins and include cats, dogs, rats, and particularly ferrets and stoats
The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
. As examples significant dog attacks have been recorded at the colony at Little Kaiteriteri Beach, and a suspected stoat
The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
or ferret
The ferret (''Mustela furo'') is a small, Domestication, domesticated species belonging to the family Mustelidae. The ferret is most likely a domesticated form of the wild European polecat (''Mustela putorius''), evidenced by their Hybrid (biol ...
attack at Doctor's Point near Dunedin
Dunedin ( ; mi, Ōtepoti) is the second-largest city in the South Island of New Zealand (after Christchurch), and the principal city of the Otago region. Its name comes from , the Scottish Gaelic name for Edinburgh, the capital of Scotland. Th ...
, New Zealand claimed the lives of 29 little blue penguins in November 2014.
Oil spills
Little penguin populations have been significantly affected by a major oil spill with the grounding of the ''Rena'' off New Zealand in 2011, which killed 2,000 seabirds (including little penguins) directly, and killed an estimated 20,000 in total based on wider ecosystem impacts.
Conservation
''Eudyptula
The genus ''Eudyptula'' ("good little diver") contains two species of penguin, found in southern Australia, Tasmania, and New Zealand (including the Chatham Islands). They are commonly known as the little penguin, little blue penguin, or, in Aus ...
'' species are classified as "at risk - declining" under New Zealand's Wildlife Act 1953
Wildlife Act 1953 is an Act of Parliament in New Zealand. Under the act, the majority of native New Zealand vertebrate species are protected by law, and may not be hunted, killed, eaten or possessed. Violations may be punished with fines of up to ...
.
Zoological exhibits
Zoological exhibits featuring purpose-built enclosures for ''Eudyptula
The genus ''Eudyptula'' ("good little diver") contains two species of penguin, found in southern Australia, Tasmania, and New Zealand (including the Chatham Islands). They are commonly known as the little penguin, little blue penguin, or, in Aus ...
'' species can be seen in Australia at the Adelaide Zoo
Adelaide Zoo is Australia's second oldest zoo (after Melbourne Zoo), and it is operated on a non-profit basis. It is located in the parklands just north of the city centre of Adelaide, South Australia. It is administered by the Royal Zoologic ...
, Melbourne Zoo
Melbourne Zoo is a zoo in Melbourne, Australia. It is located within Royal Park in Parkville, approximately north of the centre of Melbourne. It is the primary zoo serving Melbourne. The zoo contains more than 320 animal species from Austra ...
, the National Zoo & Aquarium
The National Zoo and Aquarium is a privately owned zoo and aquarium in the Australian capital city of Canberra. It is located in Yarralumla at the western end of Lake Burley Griffin, next to Scrivener Dam. It recently had a major expansion, wi ...
in Canberra
Canberra ( )
is the capital city of Australia. Founded following the federation of the colonies of Australia as the seat of government for the new nation, it is Australia's largest inland city and the eighth-largest city overall. The ci ...
, Perth Zoo
Perth Zoo is a zoological park in South Perth, Western Australia. The zoo first opened in 1898 and by 2011 housed 1258 animals of 164 species and an extensive botanical collection.
It is a full institutional member of the Zoo and Aquarium Ass ...
, Caversham Wildlife Park (Perth), Ballarat Wildlife Park
Ballarat Wildlife Park is an interactive wildlife park situated in Ballarat, Australia which was opened by Greg Parker in 1987. The Park is situated on of natural bush land.
Animals
The Park currently holds around 400 species of animals, inclu ...
, Sea Life Sydney Aquarium
SEA LIFE Sydney Aquarium (formerly Sydney Aquarium) is a public aquarium that features a large variety of Australian aquatic life, displaying more than 700 species comprising more than 13,000 individual fish and other sea and water creatures fr ...
, and the Taronga Zoo
Taronga Zoo is a zoo located in Sydney, New South Wales, Australia, in the suburb of Mosman, on the shores of Sydney Harbour. The opening hours are between 9:30 a.m. to 5 p.m. Taronga is an Aboriginal word meaning 'beautiful water view'.
It ...
in Sydney
Sydney ( ) is the capital city of the state of New South Wales, and the most populous city in both Australia and Oceania. Located on Australia's east coast, the metropolis surrounds Sydney Harbour and extends about towards the Blue Mountain ...
. Enclosures include nesting boxes or similar structures for the animals to retire into, a reconstruction of a pool and in some cases, a transparent aquarium wall to allow patrons to view the animals underwater while they swim.
''Eudyptula'' penguin exhibit exists at Sea World, on the Gold Coast, Queensland
The Gold Coast is a coastal city in the state of Queensland, Australia, approximately south-southeast of the centre of the state capital Brisbane. With a population over 600,000, the Gold Coast is the sixth-largest city in Australia, the nati ...
, Australia. In early March 2007, 25 of the 37 penguins died from an unknown toxin following a change of gravel in their enclosure. It is still not known what caused the deaths of the penguins, and it was decided not to return the 12 surviving penguins to the same enclosure where the penguins became ill. A new enclosure for the little penguin colony was opened at Sea World in 2008.
In New Zealand, ''Eudyptula'' penguin exhibits exist at the Auckland Zoo
Auckland Zoo is a zoological garden in Auckland, New Zealand, situated next to Western Springs Park not far from Auckland's central business district. It is run by the Auckland Council with the Zoological Society of Auckland as a supporting ...
, the Wellington Zoo
Wellington Zoo is a zoo in the green belt of Wellington, New Zealand.
History
Wellington Zoo was opened in 1906 by the late Prime Minister Richard Seddon after he was given a young lion – later named King Dick – by the Bostock and Wombwe ...
, and the National Aquarium of New Zealand
The National Aquarium of New Zealand, formerly Napier Aquarium, is a public aquarium on Marine Parade in Napier, New Zealand. It was started in 1957 and moved to its present location in 1976. It is owned by Napier City Council. In addition to ma ...
. Since 2017, the National Aquarium of New Zealand, has featured a monthly "Penguin of the Month" board, declaring two of their resident animals the "Naughty" and "Nice" penguin for that month. Photos of the board have gone viral and gained the aquarium a large worldwide social media following.
A colony of ''Eudyptula'' penguins exists at the New England Aquarium in Boston, Massachusetts. The penguins are one of three species on exhibit and are part of the Association of Zoos and Aquariums
The Association of Zoos and Aquariums (AZA), originally the American Association of Zoological Parks and Aquariums, is an American 501(c)(3)
nonprofit organization founded in 1924 and dedicated to the advancement of zoos and public aquariums in ...
' Species Survival Plan
The American Species Survival Plan or SSP program was developed in 1981 by the (American) Association of Zoos and Aquariums to help ensure the survival of selected species in zoos and aquariums, most of which are threatened or endangered in the wi ...
for ''Eudyptula'' penguin. ''Eudyptula'' penguins can also be seen at the Louisville Zoo
The Louisville Zoological Gardens, commonly known as the Louisville Zoo, is a zoo in Louisville, Kentucky, situated in the city's Poplar Level neighborhood. Founded in 1969, the "State Zoo of Kentucky" currently exhibits over 1,700 animals in n ...
and the Bronx Zoo
The Bronx Zoo (also historically the Bronx Zoological Park and the Bronx Zoological Gardens) is a zoo within Bronx Park in the Bronx, New York. It is one of the largest zoos in the United States by area and is the largest metropolitan zoo in ...
.
See also
Notes
References
Further reading
*
External links
State of Penguins: Little (blue) penguin – detailed and current species account of ''(Eudyptula minor)'' in New Zealand
at the International Penguin Conservation
West Coast Penguin Trust
(New Zealand)
Philip Island Nature Park website
* Gould's '' The Birds of Australia'
plate
*
{{Taxonbar, from=Q203628
little penguin
The little penguin (''Eudyptula minor'') is a species of penguin from New Zealand. They are commonly known as little blue penguins or blue penguins owing to their slate-blue plumage and are also known by their Māori name .
The Australian lit ...
little penguin
The little penguin (''Eudyptula minor'') is a species of penguin from New Zealand. They are commonly known as little blue penguins or blue penguins owing to their slate-blue plumage and are also known by their Māori name .
The Australian lit ...
Subterranean nesting birds
Birds of New Zealand
Birds described in 1781