List Of Blaptinae Genera
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Blaptinae Blaptinae is a subfamily of darkling beetles in the family Tenebrionidae Darkling beetle is the common name for members of the beetle family Tenebrionidae. The number of species in the Tenebrionidae is estimated at more than 20,000 and the fa ...
, a subfamily of darkling beetles in the family
Tenebrionidae Darkling beetle is the common name for members of the beetle family Tenebrionidae. The number of species in the Tenebrionidae is estimated at more than 20,000 and the family is cosmopolitan in distribution. Taxonomy ''Tenebrio'' is the Latin gen ...
.


Blaptinae genera

* '' Ablapsis'' Reitter, 1887 (the Palearctic) * '' Aconobius'' Casey, 1895 (North America) * '' Adamus'' Iwan, 1997 (Indomalaya) * '' Adavius'' Mulsant & Rey, 1859 (the Palearctic and Indomalaya) * '' Adoryacus'' Koch, 1963 (tropical Africa) * '' Agnaptoria'' Reitter, 1887 (the Palearctic) * '' Alaetrinus'' Iwan, 1995 (North America and the Neotropics) * '' Allophylax'' Bedel, 1906 (the Palearctic) * '' Amathobius'' Gebien, 1920 (tropical Africa) * '' Amatodes'' Dejean, 1834 (tropical Africa) * '' Amblychirus'' Koch, 1956 (tropical Africa) * '' Amblysphagus'' Fairmaire, 1896 (tropical Africa and Indomalaya) * '' Ametrocera'' Fåhraeus, 1870 (tropical Africa) * '' Ammidium'' Erichson, 1843 (tropical Africa) * '' Ammobius'' Guérin-Méneville, 1844 (the Palearctic and Indomalaya) * '' Ammodonus'' Mulsant & Rey, 1859 (North America and the Neotropics) * '' Amphithrixoides'' Bouchard & Löbl, 2008 (the Palearctic) * '' Anatrum'' Reichardt, 1936 (the Palearctic) * '' Anaxius'' Fåhraeus, 1870 (tropical Africa) * '' Anchophthalmops'' Koch, 1956 (North America and Indomalaya) * '' Anchophthalmus'' Gerstaecker, 1854 (tropical Africa) * '' Angolositus'' Koch, 1955 (tropical Africa) * '' Anomalipus'' Guérin-Méneville, 1831 (tropical Africa) * '' Apsheronellus'' Bogatchev, 1967 (the Palearctic) * '' Aptila'' Fåhraeus, 1870 (tropical Africa) * '' Arabammobius'' Grimm & Lillig, 2020 (the Palearctic and tropical Africa) * '' Asidoblaps'' Fairmaire, 1886 (the Palearctic and Indomalaya) * '' Asidodema'' Koch, 1958 (tropical Africa) * '' Asiocaedius'' G.S. Medvedev & Nepesova, 1985 (the Palearctic) * '' Atrocrates'' Koch, 1956 (tropical Africa) * '' Atrocrypticanus'' Iwan, 1999 (tropical Africa) * '' Austrocaribius'' Marcuzzi, 1954 (the Neotropics) * '' Bantodemus'' Koch, 1955 (tropical Africa) * '' Belousovia'' G.S. Medvedev, 2007 (the Palearctic) * '' Bermejoina'' Español, 1944 (the Palearctic) * '' Bioplanes'' Mulsant, 1854 (the Palearctic) * '' Bioramix'' Bates, 1879 (the Palearctic) * '' Blacodatus'' Koch, 1963 (tropical Africa) * '' Blaps'' Fabricius, 1775 (North America, tropical Africa, Indomalaya, Australasia, and Oceania) * '' Blapstinus'' Dejean, 1821 (North America, the Neotropics, and Oceania) * '' Blaptogonia'' G.S. Medvedev, 1998 (the Palearctic and Indomalaya) * '' Blastarnodes'' Koch, 1958 (tropical Africa) * '' Blenosia'' Laporte, 1840 (tropical Africa) * '' Brachyesthes'' Fairmaire, 1868 (the Palearctic and Indomalaya) * '' Brachyidium'' Fairmaire, 1883 (Indomalaya, Australasia, and Oceania) * '' Byrrhoncus'' Koch, 1954 (tropical Africa) * '' Cabirutus'' Strand, 1929 (the Palearctic and Indomalaya) * '' Caediexis'' Lebedev, 1932 (the Palearctic) * '' Caediomorpha'' Blackburn, 1888 (Australasia) * '' Caedius'' Blanchard, 1845 (the Palearctic, tropical Africa, Indomalaya, and Australasia) * '' Calaharena'' Koch, 1963 (tropical Africa) * '' Capidium'' Koch, 1954 (tropical Africa) * '' Caraboblaps'' Bauer, 1921* * '' Cenophorus'' Mulsant & Rey, 1859 (the Neotropics) * '' Clastopus'' Fairmaire, 1898 (tropical Africa) * '' Claudegirardius'' Iwan, 1999 (tropical Africa) * '' Clitobius'' Mulsant & Rey, 1859 (the Palearctic and tropical Africa) * '' Coelocnemodes'' Bates, 1879 (Indomalaya) * '' Coeloecetes'' Blair, 1929 (Indomalaya) * '' Colasia'' Koch, 1965 (the Palearctic) * '' Colophonesthes'' Koch, 1954 (tropical Africa) * '' Colpotinoides'' Kaszab, 1975 (tropical Africa) * '' Colpotinus'' Fairmaire, 1891 (the Palearctic) * '' Conibiosoma'' Casey, 1890 (North America) * '' Conibius'' LeConte, 1851 (North America, the Neotropics, and Oceania) * '' Corinta'' Koch, 1950 (tropical Africa) * '' Cornopterus'' Koch, 1950 (tropical Africa) * '' Crististibes'' Koch, 1963 (tropical Africa) * '' Crypticanus'' Fairmaire, 1897 (tropical Africa) * '' Cybotus'' Casey, 1890 (the Neotropics) * '' Cyptus'' Gerstaecker, 1871 (the Palearctic and tropical Africa) * '' Dendarophylan'' Español, 1945 (the Palearctic) * ''
Dendarus ''Dendarus'' is a genus of darkling beetles in the family Tenebrionidae. The genus is distributed from Morocco to Caucasus and exhibits a high level of diversity with 36 species, 27 of which are island endemics. Analyses of the phylogenetic relat ...
'' Dejean, 1821 (the Palearctic) * '' Diaderma'' Koch, 1960 (tropical Africa) * '' Diastolinus'' Mulsant & Rey, 1859 (the Neotropics) * '' Diestecopus'' Solier, 1848 (tropical Africa) * '' Dila'' Fischer von Waldheim, 1844 (the Palearctic) * '' Dilablaps'' Bogatchev, 1976 (the Palearctic) * '' Dilamus'' Jacquelin du Val, 1861 (the Palearctic and tropical Africa) * '' Diphyrrhynchus'' Fairmaire, 1849 (the Palearctic, tropical Africa, Indomalaya, Australasia, and Oceania) * '' Doyenus'' Iwan, 1996 (tropical Africa) * '' Drosochrus'' Erichson, 1843 (the Palearctic and tropical Africa) * '' Ectateus'' Koch, 1956 (tropical Africa) * '' Eichleria '' Kaminski, 2015 (tropical Africa) * ''
Eleodes ''Eleodes'' is a genus of darkling beetles, family Tenebrionidae. They are commonly known as pinacate beetles or desert stink beetles. They are endemic to western North America, with many species found in the Sonoran Desert. The name ''pinacate ...
'' Eschscholtz, 1829 (desert stink beetles) (North America and the Neotropics) * '' Eleodimorpha'' Blaisdell, 1909 (North America) * '' Eleoselinus'' Kaminski, 2014 (tropical Africa) * '' Embaphion'' Say, 1824 (North America) * '' Emmalus'' Erichson, 1843 (tropical Africa) * '' Ephalus'' LeConte, 1862 (North America and the Neotropics) * '' Eremostibes'' Koch, 1963 (tropical Africa) * '' Eucolus'' Mulsant & Rey, 1853 (Indomalaya) * '' Eumylada'' Reitter, 1904 (the Palearctic) * '' Eurycaulus'' Fairmaire, 1868 (the Palearctic and tropical Africa) * '' Eurynotus'' W. Kirby, 1819 (tropical Africa) * '' Eviropodus'' Koch, 1956 (tropical Africa) * '' Falsammidium'' Koch, 1960 (tropical Africa) * '' Falsocaedius'' Español, 1943 (the Palearctic) * '' Falsolobodera'' Kaszab, 1967 (the Palearctic) * '' Freyula'' Koch, 1959 (tropical Africa) * '' Glyptopteryx'' Gebien, 1910 (tropical Africa) * '' Gnaptor'' Brullé, 1831 (the Palearctic) * '' Gnaptorina'' Reitter, 1887 (the Palearctic and Indomalaya) * '' Goajiria'' Ivie & Hart, 2016 (the Neotropics) * '' Gonocephalum'' Solier, 1834 (North America, tropical Africa, Indomalaya, Australasia, and Oceania) * '' Gonopus'' Latreille, 1828 (tropical Africa) * '' Gridelliopus'' Koch, 1956 (tropical Africa) * '' Guildia'' Antoine, 1957 (the Palearctic) * '' Hadroderus'' Koch, 1956 (tropical Africa) * '' Hadrodes'' Wollaston, 1877 (the Palearctic) * '' Hadrophasis'' Ferrer, 1992 (tropical Africa) * '' Haemodus'' Gebien, 1943 (tropical Africa) * '' Hanstroemium'' Koch, 1953 (tropical Africa) * '' Helenomelas'' Ardoin, 1972 (the Palearctic) * '' Helibatus'' Mulsant & Rey, 1859 (tropical Africa) * '' Heliopates'' Dejean, 1834 (the Palearctic) * '' Heterocheira'' Dejean, 1836 (tropical Africa and Australasia) * '' Heteropsectropus'' Kaszab, 1941 (tropical Africa) * '' Heterotarsus'' Latreille, 1829 (the Palearctic, tropical Africa, and Indomalaya) * '' Hirtograbies'' Koch, 1954 (tropical Africa) * '' Holoblaps'' Bauer, 1921* * '' Hoplarion'' Mulsant & Rey, 1854 (the Palearctic) * '' Hoplitoblaps'' Fairmaire, 1889 (Indomalaya) * '' Hovademus'' Iwan, 1996 (tropical Africa) * '' Hovarygmus'' Fairmaire, 1898 (tropical Africa) * '' Hummelinckia'' Marcuzzi, 1954 (the Neotropics) * '' Isoncophallus'' Koch, 1954 (tropical Africa) * ''
Itagonia ''Itagonia'' is a genus of beetles in the family Tenebrionidae. The majority of species are found in Southwest China, but two are found in the more central and northern parts of the country, from Gansu to Hebei and Inner Mongolia Inner Mongol ...
'' Reitter, 1887 (the Palearctic) * '' Jintaium'' Ren, 1999 (the Palearctic) * '' Lariversius'' Blaisdell, 1947 (North America) * '' Lasioderus'' Mulsant & Rey, 1854 (tropical Africa) * '' Lechius'' Iwan, 1995 (tropical Africa) * '' Leichenum'' Dejean, 1834 (North America, the Palearctic, Indomalaya, Australasia, and Oceania) * '' Litoboriolus'' Español, 1945 (the Palearctic) * '' Litoborus'' Mulsant & Rey, 1854 (the Palearctic) * '' Litororus'' Reitter, 1904 (the Palearctic) * '' Loensus'' R. Lucas, 1920 (tropical Africa and Indomalaya) * '' Luebbertia'' Koch, 1963 (tropical Africa) * '' Madobalus'' Fairmaire, 1901 (tropical Africa) * '' Mateuina'' Español, 1944 (the Palearctic) * '' Medvedevia'' Chigray, 2019 (the Palearctic) * '' Medvedevoblaps'' Bouchard & Bousquet, 2021 (the Palearctic) * '' Meglyphus'' Motschulsky, 1872 (tropical Africa) * '' Melambius'' Mulsant & Rey, 1854 (the Palearctic) * '' Melanesthes'' Dejean, 1834 (the Palearctic) * '' Melanocoma'' Wollaston, 1868 (tropical Africa) * '' Melanocratus'' Fairmaire, 1895 (tropical Africa) * '' Melanopterus'' Mulsant & Rey, 1854 (tropical Africa) * '' Melansis'' Wollaston, 1864 (the Palearctic) * '' Melasmana'' Strand, 1935 (the Palearctic) * '' Menearchus'' Carter, 1920 (tropical Africa) * '' Menederes'' Solier, 1848 (tropical Africa) * '' Menederopsis'' Koch, 1954 (tropical Africa) * '' Mesomorphus'' Miedel, 1880 (the Palearctic, tropical Africa, Indomalaya, Australasia, and Oceania) * '' Messoricolum'' Koch, 1960 (tropical Africa) * '' Micrantereus'' Solier, 1848 (the Palearctic and tropical Africa) * '' Microphylacinus'' Iwan, Kaminski & Aalbu, 2011 (tropical Africa) * '' Microplatyscelis'' Kaszab, 1940 (the Palearctic) * '' Micrositus'' Mulsant & Rey, 1854 (the Palearctic) * '' Microstizopus'' Koch, 1963 (tropical Africa) * '' Minorus'' Mulsant & Rey, 1854 (tropical Africa) * '' Monodius'' Koch, 1956 (tropical Africa) * '' Montagona'' G.S. Medvedev, 1998 (the Palearctic and Indomalaya) * '' Moragacinella'' Español, 1954 (the Palearctic) * '' Myatis'' Bates, 1879 (the Palearctic) * '' Myladina'' Reitter, 1889 (the Palearctic) * '' Nalepa'' Reitter, 1887 (the Palearctic) * '' Namazopus'' Koch, 1963 (tropical Africa) * '' Nemanes'' Fairmaire, 1888 (tropical Africa) * '' Neobaphion'' Blaisdell, 1925 (North America) * '' Neoisocerus'' Bouchard, Lawrence, Davies & Newton, 2005 (the Palearctic) * '' Neopachypterus'' Bouchard, Löbl & Merkl, 2007 (the Palearctic and Indomalaya) * '' Nepalindia'' G.S. Medvedev, 1998 (Indomalaya) * '' Nesocaedius'' Kolbe, 1915 (the Palearctic and Indomalaya) * '' Nesopatrum'' Gebien, 1921 (tropical Africa) * '' Nevisia'' Marcuzzi, 1986 (the Neotropics) * ''
Nicandra ''Nicandra '' is a genus of flowering plant in the family Solanaceae, native to western South America. It was first described by Michel Adanson in 1763. The genus is named for Greek poet Nicander of Colophon, who wrote about plants – notably i ...
'' Fairmaire, 1888 (tropical Africa) * '' Nocibiotes'' Casey, 1895 (North America) * '' Notibius'' LeConte, 1851 (North America) * '' Notocorax'' Dejean, 1834 (Indomalaya) * '' Nycterinus'' Eschscholtz, 1829 (the Neotropics) * '' Ograbies'' Péringuey, 1899 (tropical Africa) * '' Oncopteryx'' Gebien, 1943 (tropical Africa) * '' Oncotus'' Blanchard, 1845 (tropical Africa) * '' Oodescelis'' Motschulsky, 1845 (the Palearctic) * '' Opatrinus'' Dejean, 1821 (the Neotropics) * '' Opatroides'' Brullé, 1832 (North America, tropical Africa, Indomalaya, and Australasia) * '' Opatrum'' Fabricius, 1775 (the Palearctic) * '' Orarabion'' Leo & Liberto, 2011 (tropical Africa) * '' Oreomelasma'' Español, 1975 (the Palearctic) * '' Otinia'' Antoine, 1942 (the Palearctic) * '' Pachymastus'' Fairmaire, 1896 (tropical Africa) * '' Parabantodemus'' Iwan, 2000 (tropical Africa) * '' Paraselinus'' Kaminski, 2013 (tropical Africa) * '' Parastizopus'' Gebien, 1938 (tropical Africa) * '' Pedinus'' Latreille, 1797 (the Palearctic and Indomalaya) * '' Penthicinus'' Reitter, 1896 (the Palearctic) * '' Penthicoides'' Fairmaire, 1896 (Indomalaya) * '' Penthicus'' Faldermann, 1836 (the Palearctic and tropical Africa) * '' Periblaps'' Bauer, 1921* * '' Periloma'' Gebien, 1938 (tropical Africa) * '' Perithrix'' Fairmaire, 1879 (the Palearctic) * '' Peyerimhoffius'' Koch, 1948 (the Palearctic) * '' Phaleriderma'' Koch, 1954 (tropical Africa) * '' Phallocentrion'' Koch, 1956 (tropical Africa) * '' Phelopatrum'' Marseul, 1876 (the Palearctic) * '' Phylacastus'' Fairmaire, 1897 (tropical Africa) * '' Phylacinus'' Fairmaire, 1896 (tropical Africa) * '' Phylan'' Sturm, 1826 (the Palearctic) * '' Phylanmania'' Ferrer, 2013 (the Palearctic) * '' Phymatoplata'' Koch, 1956 (tropical Africa) * '' Piscicula'' Robiche, 2004 (tropical Africa) * '' Planostibes'' Gemminger, 1870 (tropical Africa) * '' Platyburak'' Iwan, 1990 (Indomalaya) * '' Platyburmanicus'' Iwan, 2003 (Indomalaya) * '' Platycolpotus'' Iwan, 1997 (Indomalaya) * '' Platylus'' Mulsant & Rey, 1859 (the Neotropics) * '' Platynosum'' Mulsant & Rey, 1859 (the Palearctic) * '' Platynotoides'' Kaszab, 1975 (Indomalaya) * '' Platynotus'' Fabricius, 1801 (Indomalaya) * '' Platyprocnemis'' Español & Lindberg, 1963 (tropical Africa) * '' Platyscelis'' Latreille, 1818 (the Palearctic) * '' Plesioderes'' Mulsant & Rey, 1859 (tropical Africa and Indomalaya) * '' Pocadiopsis'' Fairmaire, 1896 (Indomalaya) * '' Pokryszkiella'' Iwan, 1996 (tropical Africa) * '' Polycoelogastridion'' Reichardt, 1936 (the Palearctic and Indomalaya) * '' Prodilamus'' Ardoin, 1969 (the Palearctic and tropical Africa) * '' Proscheimus'' Desbrochers des Loges, 1881 (the Palearctic) * '' Prosodes'' Eschscholtz, 1829 (the Palearctic and Indomalaya) * '' Protoblaps'' Bauer, 1921* * '' Psammestus'' Reichardt, 1936 (the Palearctic) * '' Psammoardoinellus'' Leo, 1981 (the Palearctic) * '' Psammogaster'' Koch, 1953 (tropical Africa) * '' Psectes'' Hesse, 1935 (tropical Africa) * '' Psectrapus'' Solier, 1848 (tropical Africa) * '' Pseudemmallus'' Koch, 1956 (tropical Africa) * '' Pseudoblaps'' Guérin-Méneville, 1834 (the Palearctic and Indomalaya) * '' Pseudognaptorina'' Kaszab, 1977 (the Palearctic and Indomalaya) * '' Pseudolamus'' Fairmaire, 1874 (the Palearctic) * '' Pseudoleichenum'' Ardoin, 1972 (tropical Africa) * '' Pseudonotocorax'' Iwan, 1997 (Indomalaya) * '' Pteroselinus'' Kaminski, 2015 (tropical Africa) * '' Pythiopus'' Koch, 1953 (tropical Africa) * '' Raynalius'' Chatanay, 1912 (tropical Africa) * '' Reichardtiellina'' Kaszab, 1982 (the Palearctic) * '' Remipedella'' Semenov, 1907 (the Palearctic) * '' Rugoplatynotus'' Kaszab, 1975 (Indomalaya) * '' Schelodontes'' Koch, 1956 (tropical Africa) * '' Schyzoschelus'' Koch, 1954 (tropical Africa) * '' Scleroides'' Fairmaire, 1883 (Australasia) * '' Scleropatroides'' Löbl & Merkl, 2003 (the Palearctic, tropical Africa, and Indomalaya) * '' Scleropatrum'' Reitter, 1887 (the Palearctic and Indomalaya) * '' Sclerum'' Dejean, 1834 (the Palearctic, tropical Africa, and Indomalaya) * '' Scymena'' Pascoe, 1866 (Australasia) * ''
Sebastianus Sebastianus (died 413), a brother of Jovinus, was an aristocrat of southern Gaul. After Jovinus usurped the throne of the western Roman Emperor Honorius in Gaul in 411, he named Sebastianus as Augustus (co-emperor) in 412. Coins bearing Sebastia ...
'' Iwan, 1996 (tropical Africa) * '' Selenepistoma'' Dejean, 1834 (tropical Africa) * '' Selinopodus'' Koch, 1956 (tropical Africa) * ''
Selinus Selinunte (; grc, Σελῑνοῦς, Selīnoûs ; la, Selīnūs , ; scn, Silinunti ) was a rich and extensive ancient Greek city on the south-western coast of Sicily in Italy. It was situated between the valleys of the Cottone and Modion ...
'' Mulsant & Rey, 1853 (tropical Africa) * '' Silvestriellum'' Koch, 1956 (tropical Africa) * '' Sinorus'' Mulsant & Revelière, 1861 (the Palearctic) * '' Sintagona'' G.S. Medvedev, 1998 (the Palearctic) * '' Sobas'' Pascoe, 1863 (Australasia) * '' Socotropatrum'' Koch, 1970 (tropical Africa) * '' Somocoelia'' Heyden & Kraatz, 1882 (the Palearctic) * '' Somocoeloplatys'' Skopin, 1968 (the Palearctic) * '' Sphaerostibes'' Koch, 1963 (tropical Africa) * '' Stenogonopus'' Gebien, 1938 (tropical Africa) * '' Stenolamus'' Gebien, 1920 (tropical Africa) * '' Stizopus'' Erichson, 1843 (tropical Africa) * '' Stridigula'' Koch, 1954 (tropical Africa) * '' Styphacus'' Fairmaire, 1901 (tropical Africa) * '' Sulpius'' Fairmaire, 1906 (tropical Africa) * '' Syntyphlus'' Koch, 1953 (tropical Africa) * '' Tagona'' Fischer von Waldheim, 1820 (the Palearctic) * '' Tagonoides'' Fairmaire, 1886 (the Palearctic) * '' Tarphiophasis'' Wollaston, 1877 (tropical Africa) * '' Thaioblaps'' Masumoto, 1989 (Indomalaya) * '' Thaumatoblaps'' Kaszab & G.S. Medvedev, 1984 (the Palearctic) * '' Tidiguinia'' Español, 1959 (the Palearctic) * '' Tonibiastes'' Casey, 1895 (North America) * '' Tonibius'' Casey, 1895 (North America) * '' Tragardhus'' Koch, 1956 (tropical Africa) * '' Trichomyatis'' Schuster, 1931 (the Palearctic) * '' Trichosternum'' Wollaston, 1861 (tropical Africa) * '' Trichoton'' Hope, 1841 (North America and the Neotropics) * '' Trigonopilus'' Fairmaire, 1893 (Indomalaya) * '' Trigonopoda'' Gebien, 1914 (Indomalaya) * '' Trigonopus'' Mulsant & Rey, 1853 (tropical Africa) * '' Trogloderus'' Leconte, 1879 (North America) * ''
Ulus Ulus may refer to: Places *Ulus, Bartın, a district in Bartin Province, Turkey *Ulus, Ankara, an important quarter in central Ankara, Turkey **Ulus (Ankara Metro), an underground station of the Ankara Metro Other uses * ''Ulus'' (newspaper), a d ...
'' Horn, 1870 (North America and the Neotropics) * '' Upembarus'' Koch, 1956 (tropical Africa) * '' Viettagona'' G.S. Medvedev & Merkl, 2003 (Indomalaya) * '' Weisea'' Semenov, 1891 (the Palearctic) * '' Wolladrus'' Iwan & Kaminski, 2016 (the Palearctic) * '' Xerolinus'' Ivie & Hart, 2016 (the Neotropics) * '' Zidalus'' Mulsant & Rey, 1853 (the Palearctic and tropical Africa) * '' Zophodes'' Fåhraeus, 1870 (tropical Africa) * '' Zoutpansbergia'' Koch, 1956 (tropical Africa) * † '' Eupachypterus'' Kirejtshuk, Nabozhenko & Nel, 2010 * † '' Palaeosclerum'' Nabozhenko & Kirejtshuk, 2017


References

{{Reflist, refs= {{Cite journal , title = Review of genus-group names in the family Tenebrionidae (Insecta, Coleoptera) , date = 2021 , last1 = Bouchard , first1 = Patrice , last2 = Bousquet , first2 = Yves , last3 = Aalbu , first3 = Rolf L. , last4 = Alonso-Zarazaga , first4 = Miguel A. , last5 = Merkl , first5 = Ottó , last6 = Davies , first6 = Anthony E. , display-authors = 4 , journal = ZooKeys , issue = 1050 , pages = 1–633 , url = https://zookeys.pensoft.net/article/64217/ , doi = 10.3897/zookeys.1050.64217 , pmid = 34385881 , pmc = 8328949 , hdl = 10261/250214 , hdl-access = free , doi-access = free *