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The crayfish snake (''Liodytes rigida''), also known commonly as the glossy crayfish snake, the glossy swampsnake, the glossy water snake, and the striped water snake, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
semiaquatic In biology, semiaquatic can refer to various types of animals that spend part of their time in water, or plants that naturally grow partially submerged in water. Examples are given below. Semiaquatic animals Semi aquatic animals include: * Ve ...
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more j ...
in the subfamily
Natricinae The Natricinae are a subfamily of colubroid snakes, sometimes referred to as a family (Natricidae). The subfamily comprises 37 genera. Members include many very common snake species, such as the European grass snakes, and the North American wa ...
of the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Colubridae. The species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to the southeastern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
, and preys mainly on crayfish.


Geographic range

''L. rigida'' is found on the
coastal plain A coastal plain is flat, low-lying land adjacent to a sea coast. A fall line commonly marks the border between a coastal plain and a piedmont area. Some of the largest coastal plains are in Alaska and the southeastern United States. The Gulf Co ...
s of both the Atlantic Coast and the
Gulf Coast The Gulf Coast of the United States, also known as the Gulf South, is the coast, coastline along the Southern United States where they meet the Gulf of Mexico. The list of U.S. states and territories by coastline, coastal states that have a shor ...
: in eastern
Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...
, southeastern Oklahoma, southern
Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the O ...
,
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...
, southern
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Miss ...
, southern
Alabama (We dare defend our rights) , anthem = "Alabama" , image_map = Alabama in United States.svg , seat = Montgomery , LargestCity = Huntsville , LargestCounty = Baldwin County , LargestMetro = Greater Birmingham , area_total_km2 = 135,765 ...
, northern
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
, southern
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
, eastern
South Carolina )'' Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...
, and southeastern
North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and ...
. There is also a disjunct population in eastern
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
.


Description

Adults of ''L. rigida'' are on average 16 inches (about 41 cm) in total length (including tail), and are heavy-bodied. The maximum recorded total length for this
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
is . ''L. rigida'' is olive brown
dorsally Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
. Additionally, two blackish dorsal stripes may or may not be present. The upper lips (labial scales) are yellow. Ventrally, it is yellow with two parallel series of black spots, which merge anteriorly into a single series. The ventral surface of the tail may have a median black line, or it may be unmarked. The dorsal scales are arranged in 19 rows at midbody. They are strongly keeled, except for the first two rows. The first row (adjacent to the
ventrals In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
) is smooth, and the second row is weakly keeled. The ventrals number 132-142. The
anal plate Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involving s ...
is divided. The
subcaudals In snakes, the subcaudal scales are the enlarged plates on the underside of the tail.Wright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . These scales may be either single or divided (pair ...
number 51-71, and are divided. Boulenger GA (1893). ''Catalogue of the Snakes in the British Museum (Natural History). Volume I., Containing the Families ... Colubridæ Aglyphæ, part.'' London: Trustees of the British Museum (Natural History). xiii + 448 pp. + Plates I-XXVIII. ("''Tropidonotus rigidus'' ", p. 240).


Diet

''L. rigida'' eats crayfish.


Reproduction

''L. rigida'' is
viviparous Among animals, viviparity is development of the embryo inside the body of the parent. This is opposed to oviparity which is a reproductive mode in which females lay developing eggs that complete their development and hatch externally from the ...
.


Subspecies

Three subspecies of ''L. rigida'' are recognized as being valid, including the
nominotypical subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
. *'' Liodytes rigida deltae'' – Delta crayfish snake *'' Liodytes rigida rigida'' – glossy crayfish snake *'' Liodytes rigida sinicola'' – Gulf crayfish snake ''
Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atten ...
'': A trinomial authority in parentheses indicates that the subspecies was originally described in a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
other than ''Liodytes''.


References


Further reading

* Behler JL, King FW (1979). ''The Audubon Society Field Guide to North American Reptiles and Amphibians''. New York: Alfred A. Knopf. 743 pp., 657 color plates. . ("Glossy Crayfish Water Snake (''Regina rigida'')", pp. 647-648 + Plate 474). * Conant R, Bridges W (1939). ''What Snake Is That? A Field Guide to the Snakes of the United States East of the Rocky Mountains''. (With 108 drawings by Edmond Malnate). New York and London: D. Appleton-Century Company. Frontispiece map + viii + 163 pp. + Plates A-C, 1-32. ("STRIPED WATER SNAKE ''Natrix rigida'' ", p. 97 + Plate 17, figure 50). * Powell R, Conant R, Collins JT (2016). ''Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America, Fourth Edition''. Boston and New York: Houghton Mifflin Harcourt. xiv + 494 pp., 47 plates, 207 figures. . ("GLOSSY SWAMPSNAKE ''Liodytes rigida'' ", p. 414 + Plate 41). * Say T (1825). "Descriptions of three new species of ''COLUBER'', inhabiting the United States". ''J. Acad. Nat. Sci. Philadelphia'' 4 (2): 237-241. (''Coluber rigidus'', new species, pp. 239–240). * Stejneger L, Barbour T (1917). ''A Check List of North American Amphibians and Reptiles''. Cambridge, Massachusetts: Harvard University Press. 125 pp. ("''Natrix rigida'' ", p. 95).


External links

* http://www.flmnh.ufl.edu/herpetology/FL-GUIDE/Reginarigida.htm {{Taxonbar, from1=Q28320463, from2=Q3014180 Liodytes Endemic reptiles of the United States Fauna of the Southeastern United States Reptiles described in 1825 Taxa named by Thomas Say