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Limacidae, also known by their common name the keelback slugs, are a taxonomic
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of medium-sized to very large, air-breathing land slugs,
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate gastropod molluscs in the superfamily Limacoidea.


Distribution

The distribution of the family Limacidae is the western Palearctic. There are 28 species of Limacidae in Russia and adjacent countries.


Anatomy

In this family, the number of haploid
chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
s lies between 21 and 25 and also lies between 31 and 35 (according to the values in this table).Barker G. M.: Gastropods on Land: ''Phylogeny, Diversity and Adaptive Morphology''. in Barker G. M. (ed.):
The biology of terrestrial molluscs
'. CABI Publishing, Oxon, UK, 2001, . 1-146, cited pages: 139 and 142.


Taxonomy


2002 taxonomy

Zhiltsov & Schileyko (2002) Zhiltsov S. S. & Schileyko A. A. (2002). "Morphology of reproductive system of ''Bielzia coerulans'' (Gastropoda, Pulmonata) and phylogenetic relations of the genus ''Bielzia''". ''Ruthenica'' 12: 73-79
abstract
elevated the subfamily Bielziinae to family level, Bielziidae, based on the morphology of the
reproductive system The reproductive system of an organism, also known as the genital system, is the biological system made up of all the anatomical organs involved in sexual reproduction. Many non-living substances such as fluids, hormones, and pheromones are als ...
of '' Bielzia coerulans''.


2005 taxonomy

The following two subfamilies were recognized in the taxonomy of Bouchet & Rocroi (2005): * subfamily Limacinae Lamarck, 1801 - synonyms: Limacopsidae Gerhardt, 1935; Bielziinae I. M. Likharev & Wiktor, 1980 * subfamily Eumilacinae I. M. Likharev & Wiktor, 1980


Genera

Genera in the family Limacidae include: subfamily Limacinae * ''
Limax ''Limax'' is a genus of air-breathing land slugs in the terrestrial pulmonate gastropod mollusk family Limacidae. The generic name ''Limax'' literally means "slug". Some species, such as the leopard slug (''L. maximus'') and the tawny garden ...
'' Linnaeus, 1758 - type genus of the family Limacidae * '' Limacus'' Lehmann, 1864; sometimes considered a subgenus of ''Limax'' * ''Bielzia'' Clessin, 1887 - with the only species '' Bielzia coerulans'' M. Bielz, 1851. Some authors, for example Russian malacologists, classify genus ''Bielzia'' within separate family Limacopsidae. * '' Caspilimax'' P. Hesse, 1926Kantor Yu I., Vinarski M. V., Schileyko A. A. & Sysoev A. V.(published online on December 22, 2009). "Catalogue of the continental mollusks of Russia and adjacent territories"
Version 2.3.
/ref> * '' Caucasolimax'' Likharev et Wiktor, 1980 * '' Gigantomilax'' O. Boettger, 1883 ** '' Gigantomilax csikii'' Soós, 1924 ** '' Gigantomilax lederi'' (Boettger, 1883) ** '' Gigantomilax majoricensis'' (Heynemann, 1863)Wiktor A., Quintana J. & Beckmann K.-H. (2007). "Redescription of "''Limax majoricensis''" (Heynemann 1863) (Gastropoda: Pulmonata: Limacidae) from the Balearic Islands. pp. 187-197. In: Beckmann K.-H.: ''Die Land- und Süßwassermollusken der Balearischen Inseln''. ConchBooks, Hackenheim, 255 pp., . * '' Lehmannia'' Heynemann, 1862 * '' Ambigolimax'' Pollonera, 1887,Marshall, B. (2015). ''Ambigolimax'' Pollonera, 1887. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=819989 on 2015-04-15; sometimes considered a subgenus of ''Lehmannia'' * '' Limacopsis'' Simroth, 1888 * '' Malacolimax'' Malm, 1868 * '' Turcomilax'' Simroth, 1901 subfamily Eumilacinae * '' Eumilax'' O. Boettger, 1881 - type genus of the subfamily Eumilacinae ** '' Eumilax brandti'' (Martens, 1880) * '' Metalimax'' Simroth, 1896


Cladogram

A
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
showing the
phylogenic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
relationships of this family to other families within the
limacoid clade The Limacoidei is a taxonomic infraorder of air-breathing land snails, semislugs and slugs, terrestrial pulmonate gastropod molluscs in the suborder Helicina Distribution The original ancestral area of limacoid families is thought to b ...
:Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". '' Journal of Biogeography'' 27(2): 379-390.
JSTOR


Ecology

Parasites of slugs in this family include
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...
e of the marsh flies
Sciomyzidae The family Sciomyzidae belongs to the typical flies (Brachycera) of the order Diptera. They are commonly called marsh flies, and in some cases snail-killing flies due to the food of their larvae. Here, the Huttoninidae, Phaeomyiidae and Tetan ...
, and others.


References


External links


Slugs of Florida
on the UF / IFAS Featured Creatures Web site {{taxonbar, from=Q1757029 pl:Pomrowcowate