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''Lilium'' () is a genus of
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
flowering plants growing from bulbs, all with large prominent flowers. They are the true lilies. Lilies are a group of flowering plants which are important in culture and literature in much of the world. Most species are native to the
northern hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
and their range is temperate climates and extends into the
subtropics The subtropical zones or subtropics are geographical and climate zones to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately 35° north and ...
. Many other plants have "lily" in their
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
s, but do not belong to the same genus and are therefore not true lilies.


Description

Lilies are tall perennials ranging in height from . They form naked or tunicless scaly underground
bulb In botany, a bulb is structurally a short stem with fleshy leaves or leaf basesBell, A.D. 1997. ''Plant form: an illustrated guide to flowering plant morphology''. Oxford University Press, Oxford, U.K. that function as food storage organs duri ...
s which are their organs of
perennation In botany, perennation is the ability of organisms, particularly plants, to survive from one germinating season to another, especially under unfavourable conditions such as drought or winter. It typically involves development of a perennating orga ...
. In some North American species the base of the bulb develops into
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
s, on which numerous small bulbs are found. Some species develop
stolon In biology, stolons (from Latin '' stolō'', genitive ''stolōnis'' – "branch"), also known as runners, are horizontal connections between organisms. They may be part of the organism, or of its skeleton; typically, animal stolons are external s ...
s. Most bulbs are buried deep in the ground, but a few species form bulbs near the soil surface. Many species form stem-roots. With these, the bulb grows naturally at some depth in the soil, and each year the new stem puts out adventitious roots above the bulb as it emerges from the soil. These roots are in addition to the basal roots that develop at the base of the bulb, a number of species also produce contractile roots that move the bulbs deeper into the soil. The flowers are large, often fragrant, and come in a wide range of colors including whites, yellows, oranges, pinks, reds and purples. Markings include spots and brush strokes. The plants are late spring- or summer-flowering. Flowers are borne in
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
s or umbels at the tip of the stem, with six tepals spreading or reflexed, to give flowers varying from funnel shape to a "Turk's cap". The tepals are free from each other, and bear a nectary at the base of each flower. The ovary is 'superior', borne above the point of attachment of the
anthers The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
. The fruit is a three-celled capsule.European Garden Flora; Volume 1 Seeds ripen in late summer. They exhibit varying and sometimes complex
germination Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
patterns, many adapted to cool temperate climates. Most cool temperate species are deciduous and dormant in winter in their native environment. But a few species native to areas with hot summers and mild winters ('' Lilium candidum'', '' Lilium catesbaei'', '' Lilium longiflorum'') lose their leaves and enter a short dormant period in summer or autumn, sprout from autumn to winter, forming dwarf stems bearing a basal rosette of leaves until, after they have received sufficient chilling, the stem begins to elongate in warming weather. The basic chromosome number is twelve (n=12).


Taxonomy

Taxonomical division in
sections Section, Sectioning or Sectioned may refer to: Arts, entertainment and media * Section (music), a complete, but not independent, musical idea * Section (typography), a subdivision, especially of a chapter, in books and documents ** Section sig ...
follows the classical division of Comber, species acceptance follows the World Checklist of Selected Plant Families, the taxonomy of section ''Pseudolirium'' is from the Flora of North America, the taxonomy of Section ''Liriotypus'' is given in consideration of Resetnik et al. 2007, the taxonomy of Chinese species (various sections) follows the Flora of China and the taxonomy of Section ''Sinomartagon'' follows Nishikawa et al. as does the taxonomy of Section ''Archelirion''. The World Checklist of Selected Plant Families, , considers ''
Nomocharis ''Nomocharis'' is a genus of flowering plants in the family Liliaceae. It consists of about 7 species native to montane regions of western China, Myanmar, and northern India. They are similar to ''Lilium'', with one of the more obvious differenc ...
'' a separate genus in its own right, however some authorities consider ''Nomocharis'' to be embedded within ''Lilium'', rather than treat it as a separate genus. The Sinomartagon are divided in 3 paraphyletic groups, while the Leucolirion are divided in 2 paraphyletic groups. There are seven sections: *''Martagon'' *''Pseudolirium'' *''Liriotypus'' *''Archelirion'' *''Sinomartagon'' *''Leucolirion'' *''Daurolirion'' There are 111 species counted in this genus. For a full list of accepted species with their native ranges, see
List of Lilium species List of ''Lilium'' species. The genus ''Lilium'' is within the tribe Lilieae of the subfamily Lilioideae, in the family Liliaceae of the order Liliales. List Species of ''Lilium'' currently accepted, with approximate native ranges, include: ...
. Some species formerly included within this genus have now been placed in other genera. These genera include ''
Cardiocrinum ''Cardiocrinum'' is a genus of bulbous plants of the lily family first described in 1846. They are native to the Himalaya, China, the Russian Far East, and Japan. The bulbs are usually formed at the soil surface. The preferred habitat is woodlan ...
'', ''
Notholirion ''Notholirion'' is a small Asian genus of bulbous plants in the lily family, Liliaceae. It is closely related to Lilium, but each individual flowers only once, and then dies after producing offsets. The bulb is covered by a tunic. Leaves are basa ...
'', ''
Nomocharis ''Nomocharis'' is a genus of flowering plants in the family Liliaceae. It consists of about 7 species native to montane regions of western China, Myanmar, and northern India. They are similar to ''Lilium'', with one of the more obvious differenc ...
'' and '' Fritillaria''.


Etymology

The botanic name ''Lilium'' is the Latin form and is a Linnaean name. The Latin name is derived from the Greek word ''leírion'', generally assumed to refer to true, white lilies as exemplified by the
Madonna lily ''Lilium candidum'', the Madonna lily or white lily, is a plant in the true lily family. It is native to the Balkans and Middle East, and naturalized in other parts of Europe, including France, Italy, and Ukraine, and in North Africa, the Canary ...
. The word was borrowed from
Coptic Coptic may refer to: Afro-Asia * Copts, an ethnoreligious group mainly in the area of modern Egypt but also in Sudan and Libya * Coptic language, a Northern Afro-Asiatic language spoken in Egypt until at least the 17th century * Coptic alphabet ...
(dial. Fayyumic) , from standard , from Demotic , from
Egyptian Egyptian describes something of, from, or related to Egypt. Egyptian or Egyptians may refer to: Nations and ethnic groups * Egyptians, a national group in North Africa ** Egyptian culture, a complex and stable culture with thousands of years of ...
"flower". Meillet maintains that both the Egyptian and the Greek word are possible loans from an extinct, substratum language of the Eastern Mediterranean. grc, κρῖνον, label=, , was used by the Greeks, albeit for lilies of any color. The term "lily" has in the past been applied to numerous flowering plants, often with only superficial resemblance to the true lily, including
water lily Water lily or water lilies may refer to: Plants * Members of family Nymphaeaceae * Formerly, members of the genus ''Nelumbo'' (the genus to which lotus belongs) * Some members of the genus ''Nymphoides'' Other uses * ''Water Lilies'' (Monet ...
, fire lily, lily of the Nile, calla lily,
trout lily ''Erythronium'', the fawn lily, trout lily, dog's-tooth violet or adder's tongue, is a genus of Eurasian and North American plants in the lily family, most closely related to tulips. The name Erythronium derives from Ancient Greek () "red" in ...
,
kaffir lily Kaffir lily is a common name which may refer to the following ornamental plants: * ''Clivia miniata'' in the family Amaryllidaceae * ''Hesperantha coccinea'' syn. ''Schizostylis coccinea'' in the family Iridaceae Iridaceae is a family of plants ...
, cobra lily, lily of the valley,
daylily A daylily or day lily is a flowering plant in the genus ''Hemerocallis'' , a member of the family Asphodelaceae, subfamily Hemerocallidoideae. Despite the common name, it is not in fact a lily. Gardening enthusiasts and horticulturists have long ...
,
ginger lily Ginger Lily is a common name for several plants and may refer to: * '' Alpinia'' * '' Hedychium'' {{Short pages monitor