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''Lilium'' () is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition ...
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
s growing from bulbs, all with large prominent flowers. They are the true lilies. Lilies are a group of flowering plants which are important in culture and literature in much of the world. Most species are native to the northern hemisphere and their range is
temperate climate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
s and extends into the subtropics. Many other plants have "lily" in their
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contra ...
s, but do not belong to the same genus and are therefore not true lilies.


Description

Lilies are tall
perennials A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
ranging in height from . They form naked or tunicless scaly underground
bulb In botany, a bulb is structurally a short stem with fleshy leaves or leaf basesBell, A.D. 1997. ''Plant form: an illustrated guide to flowering plant morphology''. Oxford University Press, Oxford, U.K. that function as food storage organs du ...
s which are their organs of
perennation In botany, perennation is the ability of organisms, particularly plants, to survive from one germinating season to another, especially under unfavourable conditions such as drought or winter. It typically involves development of a perennating organ ...
. In some North American species the base of the bulb develops into rhizomes, on which numerous small bulbs are found. Some species develop
stolon In biology, stolons (from Latin '' stolō'', genitive ''stolōnis'' – "branch"), also known as runners, are horizontal connections between organisms. They may be part of the organism, or of its skeleton; typically, animal stolons are external ...
s. Most bulbs are buried deep in the ground, but a few species form bulbs near the soil surface. Many species form stem-roots. With these, the bulb grows naturally at some depth in the soil, and each year the new stem puts out
adventitious Important structures in plant development are buds, shoots, roots, leaves, and flowers; plants produce these tissues and structures throughout their life from meristems located at the tips of organs, or between mature tissues. Thus, a living pla ...
roots above the bulb as it emerges from the soil. These roots are in addition to the basal roots that develop at the base of the bulb, a number of species also produce contractile roots that move the bulbs deeper into the soil. The flowers are large, often fragrant, and come in a wide range of colors including whites, yellows, oranges, pinks, reds and purples. Markings include spots and brush strokes. The plants are late spring- or summer-flowering. Flowers are borne in
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the sh ...
s or
umbel In botany, an umbel is an inflorescence that consists of a number of short flower stalks (called pedicels) that spread from a common point, somewhat like umbrella ribs. The word was coined in botanical usage in the 1590s, from Latin ''umbella'' " ...
s at the tip of the stem, with six
tepals A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
spreading or reflexed, to give flowers varying from funnel shape to a "Turk's cap". The tepals are free from each other, and bear a
nectary Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutuali ...
at the base of each flower. The ovary is 'superior', borne above the point of attachment of the
anthers The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
is a three-celled capsule.European Garden Flora; Volume 1
Seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s ripen in late summer. They exhibit varying and sometimes complex
germination Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, ...
patterns, many adapted to cool temperate climates. Most cool temperate species are
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, a ...
and dormant in winter in their native environment. But a few species native to areas with hot summers and mild winters (''
Lilium candidum ''Lilium candidum'', the Madonna lily or white lily, is a plant in the true lily family. It is native to the Balkans and Middle East, and naturalized in other parts of Europe, including France, Italy, and Ukraine, and in North Africa, the Canary ...
'', ''
Lilium catesbaei ''Lilium catesbaei'', sometimes known as Catesby's lily, pine lily, leopard lily, tiger lily, or southern-red lily is a native of Florida and the coastal regions of the American Southeast, where it usually grows in damp areas from Louisiana to V ...
'', ''
Lilium longiflorum ''Lilium longiflorum'' (Japanese: テッポウユリ, ''Teppōyuri''), often called the Easter lily, is a plant endemic to both Taiwan and Ryukyu Islands ( Japan). '' Lilium formosanum'', a closely related species from Taiwan, has been treated as ...
'') lose their leaves and enter a short dormant period in summer or autumn, sprout from autumn to winter, forming dwarf stems bearing a basal rosette of leaves until, after they have received sufficient chilling, the stem begins to elongate in warming weather. The basic
chromosome number Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectiv ...
is twelve (n=12).


Taxonomy

Taxonomical division in
sections Section, Sectioning or Sectioned may refer to: Arts, entertainment and media * Section (music), a complete, but not independent, musical idea * Section (typography), a subdivision, especially of a chapter, in books and documents ** Section sig ...
follows the classical division of Comber, species acceptance follows the
World Checklist of Selected Plant Families The World Checklist of Selected Plant Families (usually abbreviated to WCSP) is an "international collaborative programme that provides the latest peer reviewed and published opinions on the accepted scientific names and synonyms of selected pl ...
, the taxonomy of section ''Pseudolirium'' is from the Flora of North America, the taxonomy of Section ''Liriotypus'' is given in consideration of Resetnik et al. 2007, the taxonomy of Chinese species (various sections) follows the Flora of China and the taxonomy of Section ''Sinomartagon'' follows Nishikawa et al. as does the taxonomy of Section ''Archelirion''. The World Checklist of Selected Plant Families, , considers '' Nomocharis'' a separate genus in its own right, however some authorities consider ''Nomocharis'' to be embedded within ''Lilium'', rather than treat it as a separate genus. The Sinomartagon are divided in 3 paraphyletic groups, while the Leucolirion are divided in 2 paraphyletic groups. There are seven sections: *''Martagon'' *''Pseudolirium'' *''Liriotypus'' *''Archelirion'' *''Sinomartagon'' *''Leucolirion'' *''Daurolirion'' There are 111 species counted in this genus. For a full list of accepted species with their native ranges, see List of Lilium species. Some species formerly included within this genus have now been placed in other genera. These genera include ''
Cardiocrinum ''Cardiocrinum'' is a genus of bulbous plants of the lily family first described in 1846. They are native to the Himalaya, China, the Russian Far East, and Japan. The bulbs are usually formed at the soil surface. The preferred habitat is woodl ...
'', ''
Notholirion ''Notholirion'' is a small Asian genus of bulbous plants in the lily family, Liliaceae. It is closely related to Lilium, but each individual flowers only once, and then dies after producing offsets. The bulb is covered by a tunic. Leaves are basa ...
'', '' Nomocharis'' and ''
Fritillaria ''Fritillaria'' (fritillaries) is a genus of spring flowering herbaceous bulbous perennial plants in the lily family (Liliaceae). The type species, '' Fritillaria meleagris'', was first described in Europe in 1571, while other species from t ...
''.


Etymology

The botanic name ''Lilium'' is the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
form and is a Linnaean name. The Latin name is derived from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
word ''leírion'', generally assumed to refer to true, white lilies as exemplified by the
Madonna lily ''Lilium candidum'', the Madonna lily or white lily, is a plant in the true lily family. It is native to the Balkans and Middle East, and naturalized in other parts of Europe, including France, Italy, and Ukraine, and in North Africa, the Canary ...
. The word was borrowed from Coptic (dial. Fayyumic) , from standard , from
Demotic Demotic may refer to: * Demotic Greek, the modern vernacular form of the Greek language * Demotic (Egyptian), an ancient Egyptian script and version of the language * Chữ Nôm Chữ Nôm (, ; ) is a logographic writing system formerly used ...
, from Egyptian "flower". Meillet maintains that both the Egyptian and the Greek word are possible loans from an extinct, substratum language of the Eastern Mediterranean. grc, κρῖνον, label=, , was used by the Greeks, albeit for lilies of any color. The term "lily" has in the past been applied to numerous flowering plants, often with only superficial resemblance to the true lily, including water lily, fire lily, lily of the Nile, calla lily,
trout lily ''Erythronium'', the fawn lily, trout lily, dog's-tooth violet or adder's tongue, is a genus of Eurasian and North American plants in the lily family, most closely related to tulips. The name Erythronium derives from Ancient Greek () "red" in ...
, kaffir lily, cobra lily,
lily of the valley Lily of the valley (''Convallaria majalis'' (), sometimes written lily-of-the-valley, is a woodland flowering plant with sweetly scented, pendent, bell-shaped white flowers borne in sprays in spring. It is native throughout the cool temperate N ...
,
daylily A daylily or day lily is a flowering plant in the genus ''Hemerocallis'' , a member of the family Asphodelaceae, subfamily Hemerocallidoideae. Despite the common name, it is not in fact a lily. Gardening enthusiasts and horticulturists have long ...
,
ginger lily Ginger Lily is a common name for several plants and may refer to: * '' Alpinia'' * '' Hedychium'' {{Short pages monitor