Ligusticum Scoticum
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''Ligusticum scoticum'', known as Scots lovage, or Scottish licorice-root, is a
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
in the
celery Celery (''Apium graveolens'') is a marshland plant in the family Apiaceae that has been cultivated as a vegetable since antiquity. Celery has a long fibrous stalk tapering into leaves. Depending on location and cultivar, either its stalks, lea ...
family
Apiaceae Apiaceae or Umbelliferae is a family of mostly aromatic flowering plants named after the type genus ''Apium'' and commonly known as the celery, carrot or parsley family, or simply as umbellifers. It is the 16th-largest family of flowering plants ...
(previously Umbelliferae) found near the coasts of northern Europe and north-eastern North America. It grows up to tall and is found in rock crevices and cliff-top grassland. It is closely related to, and possibly conspecific with, ''
Ligusticum hultenii ''Ligusticum'' (lovage, licorice root) is a genus of about 60 species of flowering plants in the family Apiaceae, native to cool temperate regions of the Northern Hemisphere. Its name is believed to derive from the Italian region of Liguria.Huxl ...
'' from the coast of the northern Pacific Ocean. The plant is edible and contains the compound
sotolon Sotolon (also known as sotolone) is a lactone and an extremely powerful aroma compound, with the typical smell of fenugreek or curry at high concentrations and maple syrup, caramel, or burnt sugar at lower concentrations. Sotolon is the major arom ...
, which is also present in
fenugreek Fenugreek (; ''Trigonella foenum-graecum'') is an annual plant in the family Fabaceae, with leaves consisting of three small obovate to oblong leaflets. It is cultivated worldwide as a semiarid crop. Its seeds and leaves are common ingredients ...
. The leaves have a flavour similar to
parsley Parsley, or garden parsley (''Petroselinum crispum'') is a species of flowering plant in the family Apiaceae that is native to the central and eastern Mediterranean region (Sardinia, Lebanon, Israel, Cyprus, Turkey, southern Italy, Greece, Por ...
or celery, while the seeds taste similar to
fenugreek Fenugreek (; ''Trigonella foenum-graecum'') is an annual plant in the family Fabaceae, with leaves consisting of three small obovate to oblong leaflets. It is cultivated worldwide as a semiarid crop. Its seeds and leaves are common ingredients ...
or
cumin Cumin ( or , or Article title
) (''Cuminum cyminum'') is a


Description

''Ligusticum scoticum'' is a
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
which typically grows tall. It has triangular, twice-
ternate Ternate is a city in the Indonesian province of North Maluku and an island in the Maluku Islands. It was the ''de facto'' provincial capital of North Maluku before Sofifi on the nearby coast of Halmahera became the capital in 2010. It is off the we ...
leaves, long, with each lobe long. The edges of the leaves may be toothed, lobed or serrated, and are typically either a paler green or magenta. The
stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushro ...
branches infrequently, and bears 2–5
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s, each of which is a compound
umbel In botany, an umbel is an inflorescence that consists of a number of short flower stalks (called pedicels) that spread from a common point, somewhat like umbrella ribs. The word was coined in botanical usage in the 1590s, from Latin ''umbella'' "p ...
in diameter. There are typically 8–12 rays in both the primary and secondary umbels. Each individual flower is around in diameter and greenish-white in colour The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
are long, with five prominent ridges on each
carpel Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
. ''Ligusticum scoticum'' tastes and smells like
parsley Parsley, or garden parsley (''Petroselinum crispum'') is a species of flowering plant in the family Apiaceae that is native to the central and eastern Mediterranean region (Sardinia, Lebanon, Israel, Cyprus, Turkey, southern Italy, Greece, Por ...
or
celery Celery (''Apium graveolens'') is a marshland plant in the family Apiaceae that has been cultivated as a vegetable since antiquity. Celery has a long fibrous stalk tapering into leaves. Depending on location and cultivar, either its stalks, lea ...
.


Distribution

''Ligusticum scoticum'' is primarily an
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
plant, with a disjunct range extending from
northern Norway Northern Norway ( nb, Nord-Norge, , nn, Nord-Noreg; se, Davvi-Norga) is a geographical Regions of Norway, region of Norway, consisting of the two northernmost counties Nordland and Troms og Finnmark, in total about 35% of the Norwegian mainlan ...
to the more northerly shores of the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
, and from western
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...
to
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
. A related species, ''
Ligusticum hultenii ''Ligusticum'' (lovage, licorice root) is a genus of about 60 species of flowering plants in the family Apiaceae, native to cool temperate regions of the Northern Hemisphere. Its name is believed to derive from the Italian region of Liguria.Huxl ...
'', which was described by
Merritt Lyndon Fernald Merritt Lyndon Fernald (October 5, 1873 – September 22, 1950) was an American botanist. He was a respected scholar of the taxonomy and phytogeography of the vascular plant flora of temperate eastern North America. During his career, Fernald pub ...
in 1930 and may be better treated as a
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of ''L. scoticum'', occurs around the northern Pacific Ocean, from
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
to
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
. The southernmost occurrence of ''L. scoticum'' is at
Ballyhalbert Ballyhalbert (formerly known as Talbotston and Halbertston) is a small village and townland in County Down, Northern Ireland. It is on the east (Irish Sea) coast of the Ards Peninsula between Ballywalter and Portavogie. It is largely residential ...
in
Northern Ireland Northern Ireland ( ga, Tuaisceart Éireann ; sco, label= Ulster-Scots, Norlin Airlann) is a part of the United Kingdom, situated in the north-east of the island of Ireland, that is variously described as a country, province or region. Nort ...
.


Ecology

Within the British Isles, ''Ligusticum scoticum'' is only found on coasts where the mean July
temperature Temperature is a physical quantity that expresses quantitatively the perceptions of hotness and coldness. Temperature is measured with a thermometer. Thermometers are calibrated in various temperature scales that historically have relied o ...
is below , and this bound is likely also to apply in other parts of the species' range. Towards the southern end of its range, the plant performs poorly on south-facing sites. It grows in fissures in rocks, where it may be the only vascular plant, and also in cliff-top grassland communities dominated by ''
Festuca rubra ''Festuca rubra'' is a species of grass known by the common name red fescue or creeping red fescue. It is widespread across much of the Northern Hemisphere and can tolerate many habitats and climates. It is best adapted to well-drained soils in c ...
'' and ''
Plantago maritima ''Plantago maritima'', the sea plantain, seaside plantain or goose tongue, is a species of flowering plant in the plantain family Plantaginaceae. It has a subcosmopolitan distribution in temperate and Arctic regions, native to most of Europe, ...
''. ''Ligusticum scoticum'' cannot tolerate
grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and other ...
, and is harmed by the actions of nesting
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
s; it is therefore rarely found on bird cliffs, or where grazing sheep and rabbits are found. It is, however, tolerant of
salt spray Salt is a mineral composed primarily of sodium chloride (NaCl), a chemical compound belonging to the larger class of salts; salt in the form of a natural crystalline mineral is known as rock salt or halite. Salt is present in vast quantitie ...
, and its growth has been shown to improve when given dilute
sea water Seawater, or salt water, is water from a sea or ocean. On average, seawater in the world's oceans has a salinity of about 3.5% (35 g/L, 35 ppt, 600 mM). This means that every kilogram (roughly one liter by volume) of seawater has approx ...
. The leaves of ''L. scoticum'' are frost-tolerant, and die back each winter, but regrow very rapidly the following spring. In the British Isles,
flowering A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...
occurs from June to August, and the seeds are ripe in October or November; the timing is expected to be later at higher latitudes. The flowers of ''L. scoticum'' are visited by generalist
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...
s, mostly
flies Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...
.


Taxonomy

''Ligusticum scoticum'' was first described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his 1753 work '. Linnaeus originally used the
epithet An epithet (, ), also byname, is a descriptive term (word or phrase) known for accompanying or occurring in place of a name and having entered common usage. It has various shades of meaning when applied to seemingly real or fictitious people, di ...
', and this is used by many authors in North America; in Europe, the amended spelling ' is used.


Uses

The plant was formerly widely eaten in western Britain, both for nutrition and to combat
scurvy Scurvy is a disease resulting from a lack of vitamin C (ascorbic acid). Early symptoms of deficiency include weakness, feeling tired and sore arms and legs. Without treatment, decreased red blood cells, gum disease, changes to hair, and bleeding ...
. The leaves and stalks are edible before the flowers appear, and contain
vitamin A Vitamin A is a fat-soluble vitamin and an essential nutrient for humans. It is a group of organic compounds that includes retinol, retinal (also known as retinaldehyde), retinoic acid, and several provitamin A carotenoids (most notably bet ...
and C. They can be consumed raw or cooked like celery. Additionally, the young shoots can be candied.


References


External links

* {{Taxonbar, from=Q3257294 scoticum Flora of Northern Europe Flora of Subarctic America Flora of Eastern Canada Flora of the Northeastern United States Edible Apiaceae Plants described in 1753 Taxa named by Carl Linnaeus Flora of Saint Pierre and Miquelon Flora of Greenland