Libocedrus Tetragona
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''Pilgerodendron'' is a genus of
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
belonging to the cypress family
Cupressaceae Cupressaceae is a conifer family, the cypress family, with worldwide distribution. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdio ...
. It has only one species, ''Pilgerodendron uviferum'', which is endemic to the
Valdivian temperate rain forests The Valdivian temperate forests (NT0404) is an ecoregion on the west coast of southern South America, in Chile and Argentina. It is part of the Neotropical realm. The forests are named after the city of Valdivia. The Valdivian temperate rainforest ...
and Magellanic subpolar forests of southern Chile and southwestern Argentina. It grows from 40 to 55°S in Tierra del Fuego, where it is the southernmost conifer in the world. It is a member of subfamily Callitroideae, a group of distinct Southern Hemisphere genera associated with the Antarctic flora.Farjon, A. (2005). ''Monograph of Cupressaceae and Sciadopitys''. Royal Botanic Gardens, Kew. Flora Chilena
''Pilgerodendron uviferum''
/ref> ''Pilgerodendron'' is very closely related to the New Zealand and
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n genus ''
Libocedrus ''Libocedrus'' is a genus of five species of coniferous trees in the cypress family Cupressaceae, native to New Zealand and New Caledonia. The genus is closely related to the South American genera ''Pilgerodendron'' and '' Austrocedrus'', and ...
'', and many botanists treat ''P. uviferum'' within this genus, as ''Libocedrus uvifera'' (D.Don) Pilg.Eckenwalder, J. E. (1976). Re-evaluation of Cupressaceae and Taxodiaceae: a proposed merger. ''Madroño'' 23 (5): 237-256. It is also a taxonomical synonym for ''Libocedrus tetragona'' (Hooker). The species is known locally as ciprés de las GuaitecasChilebosque
''Pilgerodendron uviferum''
/ref> (after the Guaitecas Archipelago), and elsewhere by its scientific name, as pilgerodendron. The genus is named after Robert Knud Friedrich Pilger. ''P. uviferum'' is a
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
, slow-growing, narrowly conical evergreen tree that grows from 2–20 m in height, with a trunk up to 1.5 m in diameter. Taller specimens and wider trunks (reportedly up to 3 m in diameter) are known to have existed before it was overexploited. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are scale-like and arranged in
decussate Decussation is used in biological contexts to describe a crossing (due to the shape of the Roman numeral for ten, an uppercase 'X' (), ). In Latin anatomical terms, the form is used, e.g. . Similarly, the anatomical term chiasma is named aft ...
pairs, with each leaf equal in size, giving the shoots a square cross-section (unlike the ''Libocedrus'' species, where pairs of larger leaves alternate with pairs of smaller leaves, giving a somewhat flattened shoot). The seed cones are 5–12 mm long and 4–6 mm broad, with four scales, two sterile basal scales and two fertile scales; each scale has a slender spine-like
bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
, and each fertile scale has two winged seeds 3–4 mm long. The pollen cones are 5–10 mm long and 2 mm broad, with 12–20 scales. It is found in the evergreen coastal lowland forests along the Pacific coast of southern Patagonia, in association with the broadleaf evergreens '' Nothofagus betuloides'' and '' Drimys winteri''. It is also found in open stands in sheltered
bog A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and muskeg; a ...
s farther inland, where it is often locally dominant, and ranges as far as the eastern slopes of the Andes in southwestern Argentina. At the northern end of its range it is found in association with ''
Fitzroya cupressoides ''Fitzroya'' is a monotypic genus in the cypress family. The single living species, ''Fitzroya cupressoides'', is a tall, long-lived conifer native to the Andes mountains and coastal of southern Chile, and only to the Andes mountains Argentina, ...
''. It has also been planted along the northern part of the Pacific Coast of the United States. At present much ''Pilgerodendron uviferum'' grow in the Andes and in the
Chilean Coast Range The Chilean Coastal Range ( es, Cordillera de la Costa) is a mountain range that runs from north to south along the Pacific coast of South America parallel to the Andean Mountains, extending from Morro de Arica in the north to Taitao Peninsula, ...
. However, during the interstadials of the region's last glacial period, ''P. uviferum'' grew in lowland areas such as the Central Valley, where it is now absent. Remaining lowland populations are thought to be relicts that have survived the warmer climate of the Holocene. The wood of ''P. uviferum'' is yellow-reddish and has a distinct spicy-resinous smell, and is highly resistant to decay, which has made it very valuable as a source of timber for building construction in its native range. Much of its original lowland habitat has been cleared for this and other reasons. Listed as Vulnerable (VU A2cd v2.3) Due to over-exploitation, the species is now much scarcer than formerly, and is accordingly listed under CITES Appendix I, meaning commercial international trade is prohibited. ''P. uviferum'' is considered threatened by the World Conservation Monitoring Centre and
vulnerable Vulnerable may refer to: General * Vulnerability * Vulnerability (computing) * Vulnerable adult * Vulnerable species Music Albums * ''Vulnerable'' (Marvin Gaye album), 1997 * ''Vulnerable'' (Tricky album), 2003 * ''Vulnerable'' (The Used album) ...
by the International Union for Conservation of Nature.


History

Indigenous sea-faring nomads of the Chono and
Kawésqar The Kawésqar, also known as the Alacalufe, Kaweskar, Alacaluf or Halakwulup, are an indigenous people of South America, indigenous people who live in Chilean Patagonia, specifically in the Brunswick Peninsula, and Wellington Island, Wellington, ...
s groups used ''Pilgerodendron uviferum'' as firewood as well as wood for rows, boats and houses. During the Antonio de Vea expedition (1675–1676) Spanish explorers visited the
Guaitecas Guaitecas is Chilean commune located in Guaitecas Archipelago which are part of Aysén Province and Region. The administrative centre is Melinka, the only port and town in the wider Chonos Archipelago. Demographics According to the 2002 census of ...
and Chonos archipelagoes where the tree grew noticing the similarity to the "cypresses of Spain". Historically the wood was used to make railway sleepers, but eventually became popular for use as timber in all types of construction. In the 1860s,
Felipe Westhoff Felipe Arnold Westhoff Rodhius was a 19th-century entrepreneur known for his role in the logging of ''Pilgerodendron uviferum'' and the founding of Melinka in Guaitecas Archipelago in 1860. Westhoff was an ethnic German who migrated to South America ...
was one of various businessmen who exported ''Pilgerodendron uviferum'' north to Chilean and Peruvian ports. Westhoff, who was based in Ancud, came initially as an agent of
Ferrocarril Central Andino Ferrocarril Central Andino (FCCA) is the consortium which operates the Ferrovías Central railway in Peru linking the Pacific port of Callao and the capital Lima with Huancayo and Cerro de Pasco. As one of the Trans-Andean Railways it is the seco ...
in Peru to purchase sleepers but soon became an independent businessman. The town of
Melinka Melinka is a Chilean town in Aysén Province, Aysén Region. It is located on Ascención Island and is the administrative center of the commune of Guaitecas since 1979. The town is on a small peninsula off the main island, and shelters a small ...
was established in 1860 during the ''Pilgerodendron'' boom. This was the first permanent settlement in the Guaitecas Archipelago. After Westhoff's retirement in the 1870s,
Ciriaco Álvarez Ciriaco Álvarez was a businessman from Chonchi, Chiloé who rose to prominence in the exploitation of ''Pilgerodendron uviferum'' ( es, ciprés de las Guaitecas) in the southern Chilean archipelagoes. His dominance of the industry led him being du ...
, a native of Chonchi, rose as the most prominent ''P. uviferum'' businessman in the area, and was dubbed "The King of Pilgerodendron" ( es, El Rey del Ciprés). Many of the wooden structures in settlements of the southern Chilean archipelagos are built largely from ''P. uviferum'' wood.


References

{{Taxonbar, from=Q133103 Cupressaceae Valdivian temperate rainforest Trees of mild maritime climate Trees of subpolar oceanic climate Monotypic conifer genera Garden plants of South America Ornamental trees Dioecious plants