A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a

taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...

grouping, but rather a habit of plant growth – much like '' tree'' or ''

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

''. It comes from standard French ''liane'', itself from an Antilles French dialect word meaning to sheave.

Ecology

Lianas are characteristic of tropical moist broadleaf forests (especially seasonal forests), but may be found in temperate rainforests and temperate deciduous forests. There are also temperate lianas, for example the members of the '' Clematis'' or '' Vitis'' (wild grape) genera. Lianas can form bridges amidst the forest canopy, providing arboreal animals with paths across the forest. These bridges can protect weaker trees from strong winds. Lianas compete with forest trees for sunlight, water and nutrients from the soil. Forests without lianas grow 150% more fruit; trees with lianas have twice the probability of dying. Lianas may be found in many different plant families. One way of distinguishing lianas from trees and shrubs is based on the

stiffness Stiffness is the extent to which an object resists deformation in response to an applied force. The complementary concept is flexibility or pliability: the more flexible an object is, the less stiff it is. Calculations The stiffness, k, of a b ...

, specifically, the Young's modulus of various parts of the stem. Trees and shrubs have young twigs and smaller branches which are quite flexible and older growth such as trunks and large branches which are stiffer. A liana often has stiff young growths and older, more flexible growth at the base of the stem.

Habitat

Lianas compete intensely with trees, greatly reducing tree growth and tree reproduction, greatly increasing tree mortality, preventing tree seedlings from establishing, altering the course of regeneration in forests, and ultimately affecting tree population growth rates. Lianas also provide access routes in the forest canopy for many arboreal animals, including ants and many other invertebrates, lizards, rodents, sloths, monkeys, and lemurs. For example, in the Eastern tropical forests of Madagascar, many lemurs achieve higher mobility from the web of lianas draped amongst the vertical tree species. Many lemurs prefer trees with lianas for their roost sites. Lianas also provide support for trees when strong winds blow. However, they may be destructive in that when one tree falls, the connections made by the lianas may cause many other trees to fall. As noted by Charles Darwin, because lianas are supported by other plants, they may conserve resources that other plants must allocate to the development of structure and use them instead for growth and reproduction. In general, lianas are detrimental to the trees that support them. Growth rates are lower for trees with lianas; they directly damage hosts by mechanical abrasion and strangulation, render hosts more susceptible to ice and wind damage, and increase the probability that the host tree falls. Lianas also make the canopy of trees more accessible to animals which eat leaves. Because of these negative effects, trees which remain free of lianas are at an advantage; some species have evolved characteristics which help them avoid or shed lianas.

Examples

Some families and genera containing liana species include:

References

External links

Lianas and Climbing Plants of the Neotropics

Lianas and Climbing Plants of the Neotropics: Family Treatments
* 'Vines and Lianas' by Rhett Butler, at http://rainforests.mongabay.com/0406.htm * {{botany * Plant morphology Biology terminology Plant life-forms Plants by habit