The shagreen ray (''Leucoraja fullonica''), also known as shagreen skate or fuller's ray, is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

skate Skate or Skates may refer to: Fish *Skate (fish), several genera of fish belonging to the family Rajidae * Pygmy skates, several genera of fish belonging to the family Gurgesiellidae * Smooth skates or leg skates, several genera of fish belongin ...

in the family Rajidae. This ray is found in the eastern Atlantic Ocean, from

Murmansk Murmansk (Russian: ''Мурманск'' lit. "Norwegian coast"; Finnish: ''Murmansk'', sometimes ''Muurmanski'', previously ''Muurmanni''; Norwegian: ''Norskekysten;'' Northern Sámi: ''Murmánska;'' Kildin Sámi: ''Мурман ланнҍ'') i ...

, Russia through Norway, southern Iceland, the

Faroe Islands The Faroe Islands ( ), or simply the Faroes ( fo, Føroyar ; da, Færøerne ), are a North Atlantic island group and an autonomous territory of the Kingdom of Denmark. They are located north-northwest of Scotland, and about halfway bet ...

, the

Celtic Sea The Celtic Sea ; cy, Y Môr Celtaidd ; kw, An Mor Keltek ; br, Ar Mor Keltiek ; french: La mer Celtique is the area of the Atlantic Ocean off the southern coast of Ireland bounded to the east by Saint George's Channel; other limits includ ...

, the northern

North Sea The North Sea lies between Great Britain, Norway, Denmark, Germany, the Netherlands and Belgium. An epeiric sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian S ...

and

Skagerrak The Skagerrak (, , ) is a strait running between the Jutland peninsula of Denmark, the southeast coast of Norway and the west coast of Sweden, connecting the North Sea and the Kattegat sea area through the Danish Straits to the Baltic Sea. The ...

, to western Morocco and the

Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...

archipelago, but not in the shallow waters off

England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...

and

Wales Wales ( cy, Cymru ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by England to the Wales–England border, east, the Irish Sea to the north and west, the Celtic Sea to the south west and the ...

. It is also recorded sporadically in the

Mediterranean Sea The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the ea ...

.Guide of Mediterranean Skates and Rays (Leucoraja fullonica). Oct. 2022. Mendez L., Bacquet A. and F. Briand. http://www.ciesm.org/Guide/skatesandrays/leucoraja-fullonica

Shagreen Shagreen is a type of rawhide consisting of rough untanned skin, historically from a horse's or onager's back, or from shark or ray. Etymology The word derives from the French ''chagrin'' and is related to Italian ''zigrino'' and Venetian '' ...

is a form of roughened untanned leather, typically dyed green and commonly made of the skins of sharks and rays. The

common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...

"fuller's ray" comes from an instrument used by fullers to smooth

cloth Textile is an umbrella term that includes various fiber-based materials, including fibers, yarns, filaments, threads, different fabric types, etc. At first, the word "textiles" only referred to woven fabrics. However, weaving is not the ...

, referring to its rough and spiny back. The

pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...

disc of the shagreen ray is somewhat wider than it is long. The forward margin

concave Concave or concavity may refer to: Science and technology * Concave lens * Concave mirror Mathematics * Concave function, the negative of a convex function * Concave polygon, a polygon which is not convex * Concave set * The concavity In ca ...

and undulated, and about one-third longer than the rear margin. The

snout A snout is the protruding portion of an animal's face, consisting of its nose, mouth, and jaw. In many animals, the structure is called a muzzle, rostrum, or proboscis. The wet furless surface around the nostrils of the nose of many mammals is c ...

is cone-shaped and pointed. The

tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, r ...

is shorter than the disc, with two small, equally sized

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

s and a rudimentary

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

. The eyes and

mouth In animal anatomy, the mouth, also known as the oral cavity, or in Latin cavum oris, is the opening through which many animals take in food and issue vocal sounds. It is also the cavity lying at the upper end of the alimentary canal, bounded on ...

are large; the

teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...

are pointed and number about 64 in the

upper jaw The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...

and 56 in the

lower Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Gloucestershire, England See also *Nizhny Nizhny (russian: Ни́жний; masculine), Nizhnyaya (; feminine), or Nizhneye (russian: Ни́� ...

. The upper surface is rough in texture, with patches of large spines on the snout, around the eyes, and on the shoulder. There are two rows of about 50 spines each running from the center of the back to the first dorsal fin, and no interdorsal spine. The underside is smooth except for the front edge of the disc, base of the tail, and tail. The coloration is a uniform gray or brown with smaller darker dots above, and white below. The maximum size is to 120 cm long. This species occurs in relatively cold water on the upper

continental slope A continental margin is the outer edge of continental crust abutting oceanic crust under coastal waters. It is one of the three major zones of the ocean floor, the other two being deep-ocean basins and mid-ocean ridges. The continental margin ...

s at a depth of 30–550 m. It is most common at depths of 200 m, but is found deeper in southern areas. It favors

sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural class of s ...

y and possibly also rocky

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s. The shagreen ray feeds on a variety of

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

animal Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motilit ...

s, mainly

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

es but also

crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

s. Like other skates reproduction is

oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...

; the

egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...

capsules are amber in color and oblong-shaped, with long, stiff horns at each corner. One pair of horns are longer than the capsule. The capsules measure 7.5-9.9 cm long and 4.5-4.7 cm wide. Historically this ray was considered of "inferior" quality for consumption. It is caught

commercially Commerce is the large-scale organized system of activities, functions, procedures and institutions directly and indirectly related to the exchange (buying and selling) of goods and services among two or more parties within local, regional, nation ...

by longlines and

trawl Trawling is a method of fishing that involves pulling a fishing net through the water behind one or more boats. The net used for trawling is called a trawl. This principle requires netting bags which are towed through water to catch different speci ...

References

{{Taxonbar, from=Q383482 shagreen ray Fish of the North Sea Marine fish of Europe shagreen ray shagreen ray