Leuconotopicus Borealis
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The red-cockaded woodpecker (''Leuconotopicus borealis'') is a woodpecker endemic to the southeastern United States.


Description

The red-cockaded woodpecker is small to mid-sized species, being intermediate in size between North America's two most widespread woodpeckers (the downy and hairy woodpeckers). This species measures in length, spans across the wings and weighs . Among the standard measurements, the wing chord is , the tail is , the bill is and the tarsus is . Its back is barred with black and white horizontal stripes. The red-cockaded woodpecker's most distinguishing feature is a black cap and nape that encircle large white cheek patches. Rarely visible, except perhaps during the breeding season and periods of territorial defense, the male has a small red streak on each side of its black cap called a ''cockade'', hence its name. The species is listed as Near Threatened by the IUCN and as Endangered by the United States Fish and Wildlife Service.


Behavior

The red-cockaded woodpecker feeds primarily on ants, beetles, cockroaches,
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s, wood-boring insects, and spiders, and occasionally fruit and berries. The vast majority of foraging is on pines, with a strong preference for large trees, though they will occasionally forage on hardwoods and even on corn earworms in cornfields.Jackson, J. A. (1994). Red-cockaded Woodpecker (Dryobates borealis), version 2.0. In The Birds of North America (A. F. Poole and F. B. Gill, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bna.85 Red-cockaded woodpeckers are a territorial, nonmigratory, cooperative breeding species, frequently having the same mate for several years. The nesting season runs from April to June. The breeding female lays three to four eggs in the breeding male's roost cavity. Group members incubate the small white eggs for 10–13 days. Once hatched, the nestlings remain in the nest cavity for about 26–29 days. Upon fledging, the young often remain with the parents, forming groups of up to nine or more members, but more typically three to four members. There is only one pair of breeding birds within each group, and they normally only raise a single brood each year. The other group members, called helpers, usually males from the previous breeding season, help incubate the eggs and raise the young. Juvenile females generally leave the group before the next breeding season, in search of solitary male groups. The main predators of red-cockaded nests are rat snakes, although corn snakes also represent a threat.Longleaf Alliance
Studies have also explored the possibility that southern flying squirrels might have a negative impact on red-cockaded woodpecker populations due competition over cavities and predation on eggs and nestlings.


Distribution and habitat

Historically, this woodpecker's range extended in the southeastern United States from Florida to New Jersey and Maryland, as far west as eastern Texas and
Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...
, and inland to Missouri, Kentucky, and Tennessee. Today it is estimated that there are about 5,000 groups of red-cockaded woodpeckers, or 12,500 birds, from Florida to Virginia and west to southeast Oklahoma and eastern Texas, representing about 1% of the woodpecker's original population (over 1 million individuals at one point). They have become locally extinct ( extirpated) in Kentucky, Maryland, Missouri, New Jersey, and Tennessee. The red-cockaded woodpecker makes its home in fire-dependent pine savannas. Longleaf pines (''Pinus palustris'') are most commonly preferred, but other species of southern pine are also acceptable. While other woodpeckers bore out cavities in dead trees where the wood is rotten and soft, the red-cockaded woodpecker is the only one that excavates cavities exclusively in living pine trees. The older pines favored by the red-cockaded woodpecker often suffer from a
fungal infection Fungal infection, also known as mycosis, is disease caused by fungi. Different types are traditionally divided according to the part of the body affected; superficial, subcutaneous, and systemic. Superficial fungal infections include common ti ...
called
red heart rot Red is the color at the long wavelength end of the visible spectrum of light, next to orange and opposite violet. It has a dominant wavelength of approximately 625–740 nanometres. It is a primary color in the RGB color model and a secondary ...
which attacks the center of the trunk, causing the inner wood, the heartwood, to become soft. Cavities are generally excavated over 1 to 3 years. The aggregate of cavity trees is called a cluster and may include 1 to 20 or more cavity trees on 3 to 60
acre The acre is a unit of land area used in the imperial Imperial is that which relates to an empire, emperor, or imperialism. Imperial or The Imperial may also refer to: Places United States * Imperial, California * Imperial, Missouri * Imp ...
s (12,000 to 240,000 m²). The average cluster is about 10 acres (40,000 m²). Cavity trees that are being actively used have numerous, small resin wells which exude sap. The birds keep the sap flowing apparently as a cavity defense mechanism against rat snakes and possibly other predators. The typical territory for a group ranges from about 125 to 200 acres (500,000 to 800,000 m²), but observers have reported territories running from a low of around 60 acres (240,000 m²), to an upper extreme of more than 600 acres (2.40 km²). The size of a particular territory is related to both habitat suitability and population density. Where red-cockaded woodpeckers occur at high densities, individuals appear to spend more time in territorial defense, potentially at the expense of foraging and time allocated to reproduction, resulting in reduced clutch size and fledgling production. The red-cockaded woodpecker plays a vital role in the intricate web of life of the southern pine forests. A number of other birds and small
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s use the cavities excavated by red-cockaded woodpeckers, such as chickadees, bluebirds, titmice, and several other woodpecker species, including the downy, hairy, and
red-bellied woodpecker The red-bellied woodpecker (''Melanerpes carolinus'') is a medium-sized woodpecker of the family Picidae. It breeds mainly in the eastern United States, ranging as far south as Florida and as far north as Canada. Though it has a vivid orange-red ...
s. Larger woodpeckers such as
northern flicker The northern flicker or common flicker (''Colaptes auratus'') is a medium-sized bird of the woodpecker family. It is native to most of North America, parts of Central America, Cuba, and the Cayman Islands, and is one of the few woodpecker spec ...
, red-bellied or pileated woodpecker may take over a red-cockaded woodpecker cavity, sometimes enlarging the hole enough to allow
eastern screech owl The eastern screech owl (''Megascops asio'') or eastern screech-owl, is a small typical owl, owl that is relatively common in Eastern North America, from Mexico to Canada. This species is native to most wooded environments of its distribution, a ...
s, wood ducks, and even raccoons to move in later.
Flying squirrel Flying squirrels (scientifically known as Pteromyini or Petauristini) are a tribe of 50 species of squirrels in the family Sciuridae. Despite their name, they are not in fact capable of full flight in the same way as birds or bats, but they ar ...
s, several species of
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s and amphibians, and insects, primarily
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyly, monophyletic lineage within the ...
s and wasps, also will use red-cockaded woodpecker cavities.


Ecology

A significant competitor is the pileated woodpecker, who is known for doubling the width of cavities in trees, making the tree as a whole uninhabitable to the red-cockaded woodpecker.


Conservation

The red-cockaded woodpecker suffers from
habitat fragmentation Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological processes ...
when habitable pines are removed. When a larger cluster of birds gets split up, it is difficult for the young to find mates and eventually becomes an issue regarding
species dispersal Biological dispersal refers to both the movement of individuals (animals, plants, fungi, bacteria, etc.) from their birth site to their breeding site ('natal dispersal'), as well as the movement from one breeding site to another ('breeding dis ...
. While dispersing in search of new places to settle, the red-cockaded woodpecker encounters habitats of competing woodpecker species. The red-cockaded woodpecker has been the focus of conservation efforts even before the passing of the
Endangered Species Act The Endangered Species Act of 1973 (ESA or "The Act"; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of ec ...
in 1973. In Florida, pairs are being released at DuPuis Management Area and other privately-owned land. Due to the high importance of nesting habitat on the woodpecker's reproduction, much management has been dedicated to create ideal and more numerous nesting sites. Nesting clusters have been spared from forestry activity to preserve old-growth, large diameter trees. The nesting sites themselves have also been managed to make them more appealing. The use of controlled burning has been used to reduce deciduous growth around nesting colonies. The red-cockaded woodpecker has been shown to prefer nesting sites with less deciduous growth. The use of controlled burning must be exercised with caution due to the highly flammable resin barriers formed by the woodpecker. In an effort to increase the red-cockaded woodpecker population, states such as Texas, Louisiana, Alabama, and Georgia's wildlife management are creating artificial cavities in Longleaf Pine trees. There are two methods in which wildlife management officers use to insert cavities in long leaf pines. The most respected and latest approach is to carve out a nesting cavity in the tree and insert a man-made rot-resistant wooden box with a PVC pipe small enough for only a red-cockaded woodpecker to fit through. These boxes, also known as "inserts", can last up to 10 years. The older and less used approach is to drill a cavity into the tree in hopes that the birds will settle there and nest. Due to the energetically expensive process of excavating new cavities, more energy is expended competing for existing home ranges rather than colonizing new areas. Cavities are highly sought-out resources by all cavity dwelling species and red-cockadeds have been observed roosting in them as early as the same night the boxes are installed. Alpha males usually reside in the best cavity, alpha females in the second best cavity until breeding season when their male partners allow their cavity to turn into the nest cavity. Juveniles are left with the lower quality cavities or no cavities at all, forcing them to roost on a branch outside overnight. Due to the fact that each family member requires a cavity to roost, land managers may choose to insert additional artificial cavities to boost survival of juveniles. Red-cockaded woodpeckers will even recolonize abandoned ranges when cavities are created. In addition to the creation of new cavities, methods for protecting existing cavities are also used. The most common technique employed is a restrictor plate. The plate prevents other species from enlarging or changing the shape of the cavity entrance. These restrictor plates must be carefully monitored, however, to ensure that no hindrance is given to the woodpecker. Adjustments must also be made as the tree grows. Southern flying squirrel exclusion devices may be considered as well. A study suggested that managers establish new woodpecker clusters away from streams and limit the use of excess cavities, both factors important in the recruitment of flying squirrels. Application of capsaicin on flying squirrel at cavities could be a cost-effective method.


Intentional habitat destruction by landowners

Private landowners are prohibited from modifying habitats or taking animals that are protected by the
Endangered Species Act The Endangered Species Act of 1973 (ESA or "The Act"; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of ec ...
(ESA). In response to the listing of the Red-cockaded woodpecker, it became common in
Eastern Texas East Texas is a broadly defined cultural, geographic, and ecological region in the eastern part of the U.S. state of Texas that comprises most of 41 counties. It is primarily divided into Northeast Texas, Northeast and Southeast Texas. Most of ...
for landowners to begin cutting down the
long-leaf pine The longleaf pine (''Pinus palustris'') is a pine species native to the Southeastern United States, found along the coastal plain from East Texas to southern Virginia, extending into northern and central Florida. In this area it is also known as ...
trees so that the woodpeckers would not nest in their trees. By eliminating the trees on a piece of privately owned land, the land owners no longer have to abide by the requirements of the U.S. Fish and Wildlife Service biologists. By removing available habitat for the endangered species, landowners are able to benefit economically and maintain construction rights to the property. This led to the development of the Safe Harbor Agreements in which landowners agreed to manage their land in consistency with the conservation of a species of special conservation concern in return for relaxed ESA protection rules that allow a more flexible management of their own land.


See also

*
Babbitt v. Sweet Home Chapter of Communities for a Great Oregon ''Babbitt, Secretary of the Interior v. Sweet Home Chapter of Communities for a Great Oregon'', 515 U.S. 687 (1995), is a US Supreme Court case, decided by a 6–3 vote, in which the plaintiffs challenged the Interior Department's interpretation ...


References


Further reading

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External links


BirdLife Species Factsheet.The Nature Conservancy's Species page: Red Cockaded woodpecker
VIREO* ttp://ibc.hbw.com/ibc/phtml/especie.phtml?idEspecie=3880 Red-cockaded woodpecker videoson the Internet Bird Collection
A little bit more about the red-cockaded woodpecker and a few other endangered birds

Ecos.fws.gov
Species Profile
W.G.Jones State Forest, Conroe, TX
{{Authority control red-cockaded woodpecker Native birds of the Southeastern United States Endemic birds of the Eastern United States red-cockaded woodpecker red-cockaded woodpecker ESA endangered species