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The mountain hare (''Lepus timidus''), also known as blue hare, tundra hare, variable hare, white hare, snow hare, alpine hare, and Irish hare, is a
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores, and live solitarily or in pairs. They nest in slight depressions called forms, and their young are able to fend for themselves shortly after birth. The ge ...
that is largely adapted to polar and mountainous habitats.


Evolution

The mountain hare arose during the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division of ...
; there is evidence that its range expanded during glaciations into southern Europe, with populations of Iberian hare (''Lepus granatensis''), European hare (''L. europaeus)'' and broom hare (''L. castroviejoi'') in northern Iberia harboring
mitochondrial A mitochondrion (; ) is an organelle found in the cells of most Eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is use ...
haplotype A haplotype ( haploid genotype) is a group of alleles in an organism that are inherited together from a single parent. Many organisms contain genetic material ( DNA) which is inherited from two parents. Normally these organisms have their DNA or ...
s from the mountain hare. During the Late Pleistocene to Early Holocene, populations of mountain hare in Russia grew at least 10% larger than any living population today. This population has been classified as a distinct species ''Lepus tanaiticus'', but is now generally considered a prehistoric morphotype of the living mountain hare.


Distribution

This species is distributed from
Fennoscandia __NOTOC__ Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and no, Fennoskandia, nocat=1; russian: Фенноскандия, Fennoskandiya) or the Fennoscandian Peninsula is the geographical peninsula in Europe, which includes ...
to eastern Siberia; in addition, isolated mountain populations occur in the Alps, Scotland, the Baltics, northeastern Poland, and
Hokkaidō is Japan's second largest island and comprises the largest and northernmost prefecture, making up its own region. The Tsugaru Strait separates Hokkaidō from Honshu; the two islands are connected by the undersea railway Seikan Tunnel. The la ...
. In Ireland, the Irish hare (''L. t. hibernicus'') lives on lowland pastures, coastal grasslands, and
salt marshes A salt marsh or saltmarsh, also known as a coastal salt marsh or a tidal marsh, is a coastal ecosystem in the upper coastal intertidal zone between land and open saltwater or brackish water that is regularly flooded by the tides. It is dominated ...
, not just in the mountains. The mountain hare has also been introduced to Iceland,
Shetland Shetland, also called the Shetland Islands and formerly Zetland, is a subarctic archipelago in Scotland lying between Orkney, the Faroe Islands and Norway. It is the northernmost region of the United Kingdom. The islands lie about to the no ...
,
Orkney Orkney (; sco, Orkney; on, Orkneyjar; nrn, Orknøjar), also known as the Orkney Islands, is an archipelago in the Northern Isles of Scotland, situated off the north coast of the island of Great Britain. Orkney is 10 miles (16 km) north ...
, the Isle of Man, the Peak District,
Svalbard Svalbard ( , ), also known as Spitsbergen, or Spitzbergen, is a Norwegian archipelago in the Arctic Ocean. North of mainland Europe, it is about midway between the northern coast of Norway and the North Pole. The islands of the group range ...
,
Kerguelen Islands The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a group of islands in the sub-Antarctic constituting one of the two exposed parts of the Kerguelen Plateau, a large ...
,
Crozet Islands The Crozet Islands (french: Îles Crozet; or, officially, ''Archipel Crozet'') are a sub-Antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarcti ...
, and the Faroe Islands. In the Alps, the mountain hare lives at elevations from 700 to 3800 m, depending on geographic region and season.


Description

The mountain hare is a large species, though it is slightly smaller than the European hare. It grows to a length of , with a tail of , and a mass of , females being slightly heavier than males. They can live for up to 12 years. In summer, for all populations of mountain hares, the coat is various shades of brown. In preparation for winter most populations moult into a white (or largely white) pelage. The tail remains completely white all year round, distinguishing the mountain hare from the European hare (''Lepus europaeus''), which has a black upper side to the tail. The subspecies ''Lepus timidus hibernicus'' (the Irish mountain hare) is smaller in size and stays brown all year. The Irish hare may also have a "golden" variation, particularly those found on
Rathlin Island Rathlin Island ( ga, Reachlainn, ; Local Irish dialect: ''Reachraidh'', ; Scots: ''Racherie'') is an island and civil parish off the coast of County Antrim (of which it is part) in Northern Ireland. It is Northern Ireland's northernmost point. ...
. In the Faroe Islands, mountain hares turn grey in the winter instead of white. The winter-grey colour may be caused by downregulation of the agouti hair cycle isoform in the autumn moult.


Behaviour

The diet of the mountain hare varies from region to region. It seems to be somewhat dependent on the particular habitat in which the population under study lives. For example, in northern Scandinavia, where snow may blanket the ground for many months, the hares may feed on twigs and bark. In areas where snowfall is rare, such as Ireland, grass may form the bulk of the diet. Given a choice, mountain hares in Scotland and Ireland seem to prefer grazing (feeding on grasses). For mountain hares on a coastal grassland environment in Ireland, grasses constituted over 90% of their diet. This was higher than the percentage of grass in the diet of the European rabbit (''Oryctolagus cuniculus'') that inhabited the same environment. The mountain hare is regionally the favourite prey of the golden eagle and may additionally be preyed on by Eurasian eagle-owls and
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the Order (biology), order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe ...
es. Stoats may prey on young hares. In northern parts of Finland, Norway, and Sweden, the mountain hare and the European hare compete for habitat. The European hare, being larger, is usually able to drive away the mountain hare, but is less adapted for living in snowy regions; its feet are smaller and its winter fur is a mixture of white and brown. While this winter fur is actually a very good camouflage in the coastal regions of Finland where the snow covers the shrubs only a short time, the mountain hare is better adapted for the snowier conditions of the inland areas. The two may occasionally cross. The Arctic hare (''Lepus arcticus'') was once considered a subspecies of the mountain hare, but it is now regarded as a separate species. Similarly, some scientists believe that the Irish hare should be regarded as a separate species. Fifteen subspecies are currently recognised.


Human impact

In the European Alps, the mountain hare lives at elevations from 700 to 3800 m, depending on biographic region and season. The development of alpine winter tourism has increased rapidly since the last few decades of the 20th century, resulting in expansion of ski resorts, growing visitor numbers, and a huge increase in all forms of snow sport activities. A 2013 study looking at stress events and the response of mountain hares to disturbance concluded that those hares living in areas of high winter recreational activities showed changes in physiology and behaviour that demanded additional energy input at a time when access to food resources is restricted by snow. It recommended ensuring that forests inhabited by mountain hares were kept free of tourist development, and that new skiing areas should be avoided in mountain hare habitat, and that existing sites should not be expanded. In August 2016, the Scottish animal welfare charity OneKind launched a campaign on behalf of the mountain hare, as a way of raising awareness of mountain hare culls taking place across the country and in garnering public support for the issue. Mountain hares are routinely shot in the Scottish Highlands both as part of paid hunting "tours" and by gamekeepers managing
red grouse The red grouse (''Lagopus lagopus scotica'') is a medium-sized bird of the grouse family which is found in heather moorland in Great Britain and Ireland. It is usually classified as a subspecies of the willow ptarmigan but is sometimes consider ...
populations (who believe that mountain hares can be vectors of diseases that affect the birds). Much of this activity is secretive, but investigations have revealed that tens of thousands of hares are being culled every year. The campaign, which urges people to proclaim, "We Care For The Mountain Hare", will culminate with the charity urging the Scottish government to legislate against commercial hunting and culling of the iconic Scottish species. The campaign has revealed widespread public support for a ban on hare hunting in Scotland. On May 17, 2020, MSPs voted to ban the unlicensed culling of mountain hares and grant them protected species status within Scotland after a petition started by Green MSP Alison Johnstone gathered over 22,000 signatures. In 2021, the People's Trust for Endangered Species funded a survey of mountain hare populations in the UK's Peak District after concerns about the viability of the isolated population, believed to be as low as 2,500. The trust believes climate change is a threat to long-term survival of the Peak District population, which was introduced to the area in the 1870s.


References


External links


Irish Hare Initiative
{{Taxonbar, from=Q180035 Lepus Mammals of Asia Mammals of Europe Mammals of China Mammals of Japan Mammals of Central Asia Mammals of Russia Arctic land animals Mammals described in 1758 Taxa named by Carl Linnaeus National symbols of the Republic of Ireland Mammals of Mongolia