''Leptopelis calcaratus'' is a species of

frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...

in the family

Arthroleptidae The Arthroleptidae are a family of frogs found in sub-Saharan Africa. This group includes African treefrogs in the genus ''Leptopelis'' along with the terrestrial breeding squeakers '' Arthroleptis'', and several genera restricted to the Guinea ...

. It is found in southeastern

Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...

Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...

, the southwestern

Central African Republic The Central African Republic (CAR; ; , RCA; , or , ) is a landlocked country in Central Africa. It is bordered by Chad to the north, Sudan to the northeast, South Sudan to the southeast, the DR Congo to the south, the Republic of th ...

Equatorial Guinea Equatorial Guinea ( es, Guinea Ecuatorial; french: Guinée équatoriale; pt, Guiné Equatorial), officially the Republic of Equatorial Guinea ( es, link=no, República de Guinea Ecuatorial, french: link=no, République de Guinée équatoria ...

(including the island of

Bioko Bioko (; historically Fernando Po; bvb, Ëtulá Ëria) is an island off the west coast of Africa and the northernmost part of Equatorial Guinea. Its population was 335,048 at the 2015 census and it covers an area of . The island is located of ...

Gabon Gabon (; ; snq, Ngabu), officially the Gabonese Republic (french: République gabonaise), is a country on the west coast of Central Africa. Located on the equator, it is bordered by Equatorial Guinea to the northwest, Cameroon to the north ...

, the

Republic of the Congo The Republic of the Congo (french: République du Congo, ln, Republíki ya Kongó), also known as Congo-Brazzaville, the Congo Republic or simply either Congo or the Congo, is a country located in the western coast of Central Africa to the w ...

, and the

Democratic Republic of the Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...

. Common name Efulen forest treefrog has been coined for it (part of the

type series In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...

originated from Efulan).

Taxonomy

''Leptopelis calcaratus'' was described by the Belgian-British zoologist

George Albert Boulenger George Albert Boulenger (19 October 1858 – 23 November 1937) was a Belgian-British zoologist who described and gave scientific names to over 2,000 new animal species, chiefly fish, reptiles, and amphibians. Boulenger was also an active botani ...

in 1906 based on a specimen collected from " Efulen" in Cameroon (

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

), with additional specimens from "Cape St. John and the

Rio Benito The Benito is a river in Equatorial Guinea. It is known locally as the Mbini River, and, at least as it flows in its westerly part through the Monte Alen National Park, as the Uoro River. The river rises in Gabon and crosses into Equatorial Gui ...

District" in Equatorial Guinea; he had previously identified the latter as ''

Leptopelis rufus The red tree frog (''Leptopelis rufus'') is a species of frog in the family Arthroleptidae. It is found in southeasternmost Nigeria, western Cameroon, Equatorial Guinea, western Gabon, the lower Republic of the Congo, the lower Democratic Republi ...

''.

Subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

''Leptopelis calcaratus meridionalis'' from south of the

Congo River The Congo River ( kg, Nzâdi Kôngo, french: Fleuve Congo, pt, Rio Congo), formerly also known as the Zaire River, is the second longest river in Africa, shorter only than the Nile, as well as the second largest river in the world by discharge ...

was described by

Raymond Laurent Raymond Ferdinand Louis-Philippe Laurent (16 May 1917 – 3 February 2005) was a Belgian herpetologist, who specialized in African and South American amphibians and reptiles. He published more than 200 scientific articles and book chapters. Several ...

in 1973.

Description

Adult males measure and females in snout–vent length. This species has a characteristic white spur on the heel. The dorsum is greyish with a dark, backward-pointing triangle on the head, and a broad dark dorsal band that is often split up into bars or lateral spots. There is also often a white spot under the eye. The

canthus rostralis In snakes and amphibians, the canthus, canthal ridge or ''canthus rostralis'',Spawls S, Branch B. 1995. ''The Dangerous Snakes of Africa''. Ralph Curtis Books. Dubai: Oriental Press. 192 pp. . is the angle between the flat crown of the head and the ...

is angular. Webbing in the feet is extensive. ''L. c. meridionalis'' differs from the nominate subspecies by the spur on the heel being less developed, by having more extensive webbing, and by other small morphological differences.

Habitat and conservation

''Leptopelis calcaratus'' is an

arboreal Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. Th ...

frog found in lowland and montane rainforests at elevations from near sea level to

above sea level Height above mean sea level is a measure of the vertical distance (height, elevation or altitude) of a location in reference to a historic mean sea level taken as a vertical datum. In geodesy, it is formalized as ''orthometric heights''. The comb ...

, possibly higher. It does not survive in secondary habitats. If similar to other species of ''Leptopelis'', it would lay eggs in a nest on the ground near water. It is a common species but its habitat is affected by

habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

caused by

agricultural expansion Agricultural expansion describes the growth of agricultural land (arable land, pastures, etc.) especially in the 20th and 21st centuries. The agricultural expansion is often explained as a direct consequence of the global increase in food and ene ...

, logging, and human settlements. It occurs in a number of protected areas, including the

Korup National Park Korup National Park is in the Southwest Province of Cameroon and extends over 1,260 km2 of mostly undisturbed primary forest. It is reputedly one of Africa’s oldest and richest tropical forests in terms of floral and faunal diversity. It is ...

(Cameroon),

Monte Alén National Park Monte Alén National Park ( es, Parque nacional de Monte Alén) is located near the center of Equatorial Guinea. It was established in 1990. With an area of , it is the country's largest national park. The goliath frog ''(Conraua goliath)'', on ...

(Equatorial Guinea), and

Dzanga-Ndoki National Park The Dzanga-Ndoki National Park is located in the southwestern extremity of the Central African Republic. Established in 1990, the national park is . The national park is split into two non-continuous sectors, the northern Dzanga sector (or Dzanga ...

(Central African Republic).

References

{{Taxonbar, from=Q1947233 calcaratus Frogs of Africa Amphibians of West Africa Amphibians of Cameroon Amphibians of the Central African Republic Amphibians of the Democratic Republic of the Congo Amphibians of Equatorial Guinea Amphibians of Gabon Fauna of Nigeria Amphibians of the Republic of the Congo Amphibians described in 1906 Taxa named by George Albert Boulenger Taxonomy articles created by Polbot