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''Leptobrachium boringii'', commonly known as the Emei moustache toad or Taosze spiny toad, is a species of 
amphibian
Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...
in the family Megophryidae
Megophryidae, commonly known as goose frogs, is a large family of frogs native to the warm southeast of Asia, from the Himalayan foothills eastwards, south to Indonesia and the Greater Sunda Islands in Maritime Southeast Asia, and extending to th ...
. It is endemic
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to China
China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
where it is found in Sichuan
Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
, Guizhou
Guizhou (; formerly Kweichow) is a landlocked province in the southwest region of the People's Republic of China. Its capital and largest city is Guiyang, in the center of the province. Guizhou borders the autonomous region of Guangxi to t ...
, and Hunan
Hunan (, ; ) is a landlocked province of the People's Republic of China, part of the South Central China region. Located in the middle reaches of the Yangtze watershed, it borders the province-level divisions of Hubei to the north, Jiangxi to ...
provinces. "Emei" or "Taosze" in its common names refer to its type locality, Taosze on Mount Emei
Mount Emei (; ), alternately Mount Omei, is a mountain in Sichuan Province, China, and is the highest of the Four Sacred Buddhist Mountains of China. Mount Emei sits at the western rim of the Sichuan Basin. The mountains west of it are know ...
, Sichuan. Its natural habitat
In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s are temperate forest
A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s, grassland
A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
, arable land
Arable land (from the la, arabilis, "able to be ploughed") is any land capable of being ploughed and used to grow crops.''Oxford English Dictionary'', "arable, ''adj''. and ''n.''" Oxford University Press (Oxford), 2013. Alternatively, for the ...
, and rural gardens near river
A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of wate ...
s. It is threatened by habitat loss
Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
.
Reproductive behaviour and strategies
Male ''Leptobrachium boringii'' exhibit conspicuous keratinized nuptial spines that grow on their upper lip during the breeding season—these are the "moustache" and "spines" referred to in its common names. Moreover, male ''Leptobrachium boringii'' are larger, on average in snout-vent length, than females, which are on average in SVL (female-biasedsexual size dimorphism
Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
is more common in frogs). These unusual features seem to relate to male–male combat for best breeding territories and/or female preference for larger males. However, also multiple paternity could be observed, suggesting that some males use the sneaker strategy to fertilize eggs, instead of defending territories.
References
{{Taxonbar, from=Q28036831 boringii Amphibians of China Endemic fauna of China Taxonomy articles created by Polbot Amphibians described in 1945 Endangered Fauna of China