Leptacme Cuongi
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''Leptacme cuongi'' is a species of air-breathing
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
, a terrestrial
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk in the family Clausiliidae, the door snails. This species is known to occur in a limestone area in Thanh Hóa Province, Vietnam. The shell is high- spired and slender with numerous whorls, and is usually a little over 10 mm in length. When the species was described in 2007, as well as a detailed shell description, full accounts of the reproductive system and the
radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...
of this species were also given. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''cuongi'' is in honor of the biologist Duong Ngoc Cuong, who works at the
Institute of Ecology and Biological Resources An institute is an organisational body created for a certain purpose. They are often research organisations (research institutes) created to do research on specific topics, or can also be a professional body. In some countries, institutes can ...
in Hanoi, Vietnam, and who is a friend of the first author.


Distribution

This species occurs in: * Vietnam The type locality is Pu Luong Nature Reserve, a limestone hill near the native village Am, 20°27.39'N 105°13.65'E, in Thanh Hóa Province, Vietnam.


Description

The shell is
dextral Sinistral and dextral, in some scientific fields, are the two types of chirality ("handedness") or relative direction. The terms are derived from the Latin words for "left" (''sinister'') and "right" (''dexter''). Other disciplines use different ...
, not decollate, light corneous, fragile, glossy and rather translucent. It is very slender, with a nearly cylindrical, narrow, upper third and a spindle-shaped lower part. The shell has 12-15 moderately convex whorls separated by an indented, relatively deep suture.
Protoconch A protoconch (meaning first or earliest or original shell) is an embryonic or larval shell which occurs in some classes of molluscs, e.g., the initial chamber of an ammonite or the larval shell of a gastropod. In older texts it is also called ...
is glossy, teleoconch whorls are with obtuse growth-lines, which are more rib-like on the more narrowly sculptured, cervical part of the last whorl. Peristome is double; inner lip recognizable as a narrow ridge, contacting the body whorl and closing the umbilicus (visible in not fully grown specimens); outer lip very strongly thickened, with some prominent growth-ridges, tube-like protruding. Neck is with a more or less clearly developed basal crest. Aperture is obliquely pear-shaped to narrowed elliptical, with a sinulus bordered by the lamella parietalis and a knob on the peristome, whitish inside. Peristome is continuous, white, broadly reflexed. In frontal view, the prominent lamella parietalis (= superior), and the more inconspicuous lamella columellaris (= inferior) and plica principalis are seen, whereas the frontal end of the lamella subcolumellaris is usually discernible as a blunt denticle. The lamella parietalis is connected with the spiralis, which reaches as far as the lamella subcolumellaris inside, i.e. hardly further than the columellaris; there is no lamella inserta. Ventro-laterally, next to the plica principalis, there are four short, nearly parallel plicae palatales. Clausilial blade is rather broad, curvature of its outer border dictated by the position of the short palatal plicae. The width of the shell is 2.0–2.6 mm, 2.2 mm on average. The height of the shell is 10.4 mm (12 whorls)–14.0 mm (15 whorls), 11.9 mm on average. The conchologically most similar species, '' Leptacme eregia'' Szekeres, 1969, can be distinguished from ''Leptacme cuongi'' by its much larger shell (17.8 mm instead of 11.9 mm), its more rounded neck, i.e. without a basal crest, the lamella inferior which is situated more deeply inside, and the plica principalis, which ends less deep. Reproductive system: Vagina is more than twice the length of the oviduct. Oviduct about half as long as the proximal part of the pedunculus, which is longer than the distal part. Basally the pedunculus is much broader than the adjoining oviduct. The bursa of the bursa copulatrix is separate from the spermoviduct; it reaches hardly further than the diverticulum. Male part of the genitalia with a single retractor muscle, without flagellum or caecum. Penis is tripartite as far as can be judged on the basis of the transparent genital slides, forming a loop with a sharp curvature about halfway. Proximal half formed by (1) a short segment with a relatively simple luminal surface, followed by (2) a clearly broader and about three times longer part with some longitudinal ridges with zigzag borders. Distal half, with the insertion of the retractor muscle slightly distal from halfway, about as broad or little broader than the narrow part of the proximal half; penial segment (3) with a relatively simple luminal surface; epiphallus (4) with a vague pattern of small pustulae, most clearly seen at its distal end. Vas deferens, a short part of the penis and nearly as much as the proximal half of the vagina united by a common sheath. The large main cusp of the central tooth of
radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...
is accompanied by prominent ectocones. The 7-8 adjoining lateral teeth in half a row have equally small ectocones. All these teeth have prominent basal plates with interrow support ridges. In the first marginal tooth, the main cusp is less symmetrical and the basal plate is less prominently developed. The following c.10 marginal teeth are increasingly more asymmetrical initially, with an endocone on the main cusp and ectones with two or three cusps in the teeth closer to the margin, which are irregularly comb-like when the main cusp with the endocone becomes relatively smaller.


Ecology

''Leptacme cuongi'' is known to inhabit limestone areas.


References

This article incorporates CC-BY-3.0 text from the reference {{Taxonbar, from=Q6527862 Clausiliidae Gastropods described in 2007