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''Pistacia lentiscus'' (also lentisk or mastic) is a
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
evergreen shrub or small tree of the genus ''
Pistacia ''Pistacia'' is a genus of flowering plants in the cashew family, Anacardiaceae. It contains 10 to 20 species that are native to Africa and Eurasia from the Canary Islands, all of Africa, and southern Europe, warm and semidesert areas across Asi ...
'' native to the
Mediterranean Basin In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and w ...
. It grows up to tall and is cultivated for its aromatic resin, mainly on the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
island of
Chios Chios (; el, Χίος, Chíos , traditionally known as Scio in English) is the fifth largest Greek island, situated in the northern Aegean Sea. The island is separated from Turkey by the Chios Strait. Chios is notable for its exports of mastic ...
and around the
Turkish Turkish may refer to: *a Turkic language spoken by the Turks * of or about Turkey ** Turkish language *** Turkish alphabet ** Turkish people, a Turkic ethnic group and nation *** Turkish citizen, a citizen of Turkey *** Turkish communities and mi ...
town of
Çeşme Çeşme () is a coastal town and the administrative centre of the district of the same name in Turkey's westernmost end, on a promontory on the tip of the peninsula that also carries the same name and that extends inland to form a whole with the ...
.''Pistacia lentiscus'' L.
a
Mansfeld's Database Taxonomy


Description

The plant is evergreen, from 1 to 5 m high, with a strong smell of
resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on natu ...
, growing in dry and rocky areas in North Africa and
Mediterranean Europe Southern Europe is the southern region of Europe. It is also known as Mediterranean Europe, as its geography is essentially marked by the Mediterranean Sea. Definitions of Southern Europe include some or all of these countries and regions: Alba ...
. It resists mild to heavy frosts but prefers milder winters and grows on all types of soils, and can grow well in limestone areas and even in salty or saline environments, making it more abundant near the sea. It is also found in woodlands,
dehesa A ''dehesa'' () is a multifunctional, agrosylvopastoral system (a type of agroforestry) and cultural landscape of southern and central Spain and southern Portugal; in Portugal, it is known as a ''montado''. Its name comes from the Latin 'defens ...
s (almost deforested pasture areas),
Kermes oak ''Quercus coccifera'', the kermes oak, is an oak bush in the ''Ilex'' section of the genus. It is native to the Mediterranean region and Northern African Maghreb, south to north from Morocco to France and west to east from Portugal to Cyprus an ...
woods, wooded areas dominated by other oaks, garrigues, maquis shrublands, hills, gorges, canyons, and rocky hillsides of the entire Mediterranean area. It is a typical species of Mediterranean mixed communities which include myrtle, Kermes oak, Mediterranean dwarf palm,
buckthorn ''Rhamnus'' is a genus of about 110 accepted species of shrubs or small trees, commonly known as buckthorns, in the family Rhamnaceae. Its species range from tall (rarely to ) and are native mainly in east Asia and North America, but found thr ...
and
sarsaparilla Sarsaparilla often refers to the sarsaparilla soft drink, made from Smilax plants. Sarsaparilla may also refer to: Biology *Several species of plants, of the genus ''Smilax'', including: **''Smilax ornata'', also known as Honduran or Jamaican sar ...
, and serves as protection and food for birds and other fauna in this ecosystem. It is a very hardy pioneer species dispersed by birds. When older, it develops some large trunks and numerous thicker and longer branches. In appropriate areas, when allowed to grow freely and age, it often becomes a tree of up to 7 m. However, logging, grazing, and fires often prevent its development. The leaves are alternate, leathery, and compound paripinnate (no terminal leaflet) with five or six pairs of deep-green leaflets. It presents very small flowers, the male with five
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s, the female with a 3-part style. The fruit is a
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
, first red and then black when ripe, about 4 mm in diameter. The fruit, although not commonly consumed, is edible and has a tart raisin-like flavour. ''Pistacia lentiscus'' is related to ''
Pistacia terebinthus ''Pistacia terebinthus'' also called the terebinth and the turpentine tree, is a deciduous tree species of the genus ''Pistacia'', native to the Mediterranean region from the western regions of Morocco and Portugal to Greece and western and s ...
'', with which it hybridizes frequently in contact zones. ''Pistacia terebinthus'' is more abundant in the mountains and inland and the mastic is usually found more frequently in areas where the Mediterranean influence of the sea moderates the climate. The mastic tree does not reach the size of the ''Pistacia terebinthus'', but the hybrids are very difficult to distinguish. The mastic has winged stalks to its leaflets, i.e., the stalks are flattened and with side fins, whereas these stems in ''Pistacia terebinthus'' are simple. On the west coast of the Mediterranean,
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
and
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabian Peninsula, Arabia (including the Arabian Peninsula and Bahrain), Anatolia, Asia Minor (Asian part of Turkey except Hatay Pro ...
, it can be confused with '' P. atlantica''.


Distribution

''Pistacia lentiscus'' is native throughout the Mediterranean region, from Morocco and the
Iberian peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
in the west through southern France and Turkey to Iraq and Iran in the east. It is also native to the Canary Islands.


Ornamental use

In urban areas near the sea, where "palmitos" or Mediterranean dwarf palms grow, and other exotic plants, it is often used in gardens and resorts, because of its strength and attractive appearance. Unlike other species of ''
Pistacia ''Pistacia'' is a genus of flowering plants in the cashew family, Anacardiaceae. It contains 10 to 20 species that are native to Africa and Eurasia from the Canary Islands, all of Africa, and southern Europe, warm and semidesert areas across Asi ...
'', it retains its leaves throughout the year. It has been introduced as an ornamental shrub in
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
, where it has naturalized and is often seen primarily in suburban and semiarid areas where the summer rainfall climate, contrary to the Mediterranean, does not affect it.


Resin

The aromatic, ivory-coloured resin, also known as
mastic Mastic may refer to: Adhesives and pastes *Mastic (plant resin) *Mastic asphalt, or asphalt, is a sticky, black and highly viscous liquid *Mastic cold porcelain, or salt ceramic, is a traditional salt-based modeling clay. *Mastic, high-grade cons ...
, is harvested as a spice from the cultivated mastic trees grown in the south of the Greek island of Chios in the Aegean Sea, where it is also known by the name "Chios tears". Originally liquid, it is hardened, when the weather turns cold, into drops or patties of hard, brittle, translucent resin. When chewed, the resin softens and becomes a bright white and
opaque Opacity or opaque may refer to: * Impediments to (especially, visible) light: ** Opacities, absorption coefficients ** Opacity (optics), property or degree of blocking the transmission of light * Metaphors derived from literal optics: ** In lingu ...
gum. The word mastic derives from the Latin word ''masticare'' (to chew), in Greek: μαστιχάω verb ''mastichein'' ("to gnash the teeth", the English word completely from the Latin
masticate Chewing or mastication is the process by which food is crushed and ground by teeth. It is the first step of digestion, and it increases the surface area of foods to allow a more efficient break down by enzymes. During the mastication process, t ...
) or ''massein'' ("to chew").Mastic
a
e-xios.gr Chios Portal
/ref> Within the European Union,
mastic Mastic may refer to: Adhesives and pastes *Mastic (plant resin) *Mastic asphalt, or asphalt, is a sticky, black and highly viscous liquid *Mastic cold porcelain, or salt ceramic, is a traditional salt-based modeling clay. *Mastic, high-grade cons ...
production in
Chios Chios (; el, Χίος, Chíos , traditionally known as Scio in English) is the fifth largest Greek island, situated in the northern Aegean Sea. The island is separated from Turkey by the Chios Strait. Chios is notable for its exports of mastic ...
is granted
protected designation of origin The protected designation of origin (PDO) is a type of geographical indication of the European Union and the United Kingdom aimed at preserving the designations of origin of food-related products. The designation was created in 1992 and its main ...
and
protected geographical indication Three European Union schemes of geographical indications and traditional specialties, known as protected designation of origin (PDO), protected geographical indication (PGI), and traditional specialities guaranteed (TSG), promote and protect nam ...
names. Although the tree is native to all of the Mediterranean region, it will release its resin only on selected places, most notably, around Cesme, Turkey and in the southern portion of the Greek island of Chios, the latter being the only place in the world where it is cultivated regularly. The island's mastic production is controlled by a
co-operative A cooperative (also known as co-operative, co-op, or coop) is "an autonomous association of persons united voluntarily to meet their common economic, social and cultural needs and aspirations through a jointly owned and democratically-control ...
of "medieval" villages, collectively known as the '
mastichochoria Mastichochoria ( el, Μαστιχοχώρια, , lit. "the mastic villages" in English) is a former municipality on the island of Chios (Χίος), North Aegean, Greece. Since the 2011 local government reform it is part of the municipality Chios, ...
' (Μαστιχοχώρια, lit. "mastic villages").


Cultivation history

The resin is collected by bleeding the trees from small cuts made in the bark of the main branches, and allowing the sap to drip onto the specially prepared ground below. The harvesting is done during the summer between June and September. After the mastic is collected, it is washed manually and is set aside to dry, away from the sun, as it will start melting again. Mastic resin is a relatively expensive kind of spice; it has been used principally as a chewing gum for at least 2,400 years. The flavour can be described as a strong, slightly smoky, resiny aroma and can be an
acquired taste An acquired taste is an appreciation for something unlikely to be enjoyed by a person who has not had substantial exposure to it. It is the opposite of innate taste, which is the appreciation for things that are enjoyable by most persons without ...
. Some scholars identify the ''bakha'' בכא mentioned in the Bible—as in the Valley of Baca ( he, עמק הבכא) of Psalm 84—with the mastic plant. The word ''bakha'' appears to be derived from the Hebrew word for crying or weeping, and is thought to refer to the "tears" of resin secreted by the mastic plant, along with a sad weeping noise which occurs when the plant is walked on and branches are broken. The Valley of Baca is thought to be a valley near Jerusalem that was covered with low mastic shrubbery, much like some hillsides in northern Israel today. In an additional biblical reference, King David receives divine counsel to place himself opposite the
Philistines The Philistines ( he, פְּלִשְׁתִּים, Pəlīštīm; Koine Greek (LXX): Φυλιστιείμ, romanized: ''Phulistieím'') were an ancient people who lived on the south coast of Canaan from the 12th century BC until 604 BC, when ...
coming up the
Valley of Rephaim The Valley of Rephaim ( he, עמק רפאים, ''Emeq Rephaim'') (; , R.V.) or Valley of the Rephaim,Jerusalem Bible (1966) adds "the": 1 Chronicles 14:9 is a valley descending southwest from Jerusalem to Nahal Sorek below, it is an ancient route fr ...
, southwest of Jerusalem, such that the "sound of walking on the tops of the ''bakha'' shrubs" (קול צעדה בראשי הבכאים) signals the moment to attack (II Samuel V: 22–24). Mastic is known to have been popular in Roman times when children chewed it, and in medieval times, it was highly prized for the sultan's harem both as a breath freshener and for cosmetics. It was the sultan's privilege to chew mastic, and it was considered to have healing properties. The spice's use was widened when Chios became part of the
Ottoman Empire The Ottoman Empire, * ; is an archaic version. The definite article forms and were synonymous * and el, Оθωμανική Αυτοκρατορία, Othōmanikē Avtokratoria, label=none * info page on book at Martin Luther University) ...
, and it remains popular in North Africa and the Near East. An unflattering reference to mastic-chewing was made in Shakespeare's ''Troilus and Cressida'' (published 1609) when Agamemnon dismisses the views of the cynic and satirist Thersites as graceless productions of "his mastic jaws".


Culinary use

Mastic gum is principally used either as a flavouring or for its gum properties, as in mastic chewing gum. As a spice, it continues to be used in Greece to flavour spirits and liqueurs (such as Chios's native drink ''
mastiha Mastika or mastiha is a liqueur seasoned with mastic, a resin with a slightly pine or cedar-like flavor gathered from the mastic tree, a small evergreen tree native to the Mediterranean region. In Greece, mastiha ( el, μαστίχα) or masti ...
''), chewing gum, and a number of cakes, pastries,
spoon sweets A spoon is a utensil consisting of a shallow bowl (also known as a head), oval or round, at the end of a handle. A type of cutlery (sometimes called flatware in the United States), especially as part of a table setting, place setting, it is used ...
, and desserts. Sometimes, it is even used in making cheese."The Magic Tree — Marvelous Masticha", Epikouria Magazine, Fall/Winter 2005 Mastic resin is a key ingredient in ''
dondurma Dondurma is the common word in Turkish for all kinds of ice cream that is often used to refer to Turkish mastic ice cream in English. It typically includes the ingredients cream, whipped cream, salep (ground-up tuber of an orchid), mastic (plant ...
'' and Turkish puddings, giving those confections their unusual texture and bright whiteness. In Lebanon and Egypt, the spice is used to flavour many dishes, ranging from soups to meats to desserts, while in Morocco, smoke from the resin is used to flavour water. In Turkey, mastic is used as a flavor of
Turkish delight Turkish delight or lokum ( ota, لوقوم) is a family of confections based on a gel of starch and sugar. Premium varieties consist largely of chopped dates, pistachios, hazelnuts or walnuts bound by the gel; traditional varieties are often f ...
. Recently, a mastic-flavoured fizzy drink has also been launched, called "Mast". Mastic resin is a key ingredient in Greek festival breads, for example, the sweet bread ''tsoureki'' and the traditional New Year's ''
vasilopita Vasilopita ( el, Βασιλόπιτα, ''Vasilópita'', lit. '(St.) Basil-pie' or 'Vassilis pie', see below) is a New Year's Day bread or cake in Greece and many other areas in eastern Europe and the Balkans which contains a hidden coin or trink ...
''. Furthermore, mastic is also essential to ''
myron Myron of Eleutherae ( grc, Μύρων, ''Myrōn'' ), working c. 480–440 BC, was an Athenian sculptor from the mid-5th century BC. He was born in Eleutherae on the borders of Boeotia and Attica. According to Pliny's '' Natural History'', Agelad ...
'', the holy oil used for
chrismation Chrismation consists of the sacrament or mystery in the Eastern Orthodox, Oriental Orthodox and Eastern Catholic churches, as well as in the Assyrian Church of the East initiation rites. The sacrament is more commonly known in the West as confirm ...
by the Orthodox Churches. Mastic continues to be used for its gum and medicinal properties, as well as its culinary uses. Jordanian
chewing gum Chewing gum is a soft, cohesive substance designed to be chewed without being swallowed. Modern chewing gum is composed of gum base, sweeteners, softeners/plasticizers, flavors, colors, and, typically, a hard or powdered polyol coating. Its te ...
manufacturer, Sharawi Bros., use the mastic of this shrub as a primary ingredient in their mastic-flavoured products and they distribute the gum to many
deli Deli may refer to: * Delicatessen, a shop selling specially prepared food, or food prepared by such a shop * Sultanate of Deli, a former sultanate in North Sumatra, Indonesia Places * Deli, Boyer-Ahmad, a village in Kohgiluyeh and Boyer-Ahmad Pro ...
stores worldwide. The resin is used as a primary ingredient in the production of cosmetics such as toothpaste, lotions for the hair and skin, and perfumes.


Medicine

People in the Mediterranean region have used mastic as a medicine for gastrointestinal ailments for several thousand years. First-century Greek physician and botanist
Dioscorides Pedanius Dioscorides ( grc-gre, Πεδάνιος Διοσκουρίδης, ; 40–90 AD), “the father of pharmacognosy”, was a Greek physician, pharmacologist, botanist, and author of ''De materia medica'' (, On Medical Material) —a 5-vol ...
wrote about the medicinal properties of mastic in his classic treatise ''De Materia Medica'' (''About Medical Substances''). Some centuries later, Markellos Empeirikos and Pavlos Eginitis also noticed the effect of mastic on the digestive system. Mastic oil has antibacterial and antifungal properties, and as such is widely used in the preparation of ointments for skin disorders and afflictions. It is also used in the manufacture of plasters. In recent years, university researchers have provided the scientific evidence for the medicinal properties of mastic. A 1985 study by the
University of Thessaloniki A university () is an institution of higher (or tertiary) education and research which awards academic degrees in several academic disciplines. Universities typically offer both undergraduate and postgraduate programs. In the United States, the ...
and by the
Meikai University is a private university in Urayasu, Chiba, Japan. History The university was founded in 1970, and obtained its present name in 1988. Programs The university's School of Dentistry is located in Sakado, Saitama is a List of cities in Japan ...
discovered that mastic can reduce bacterial dental plaque in the mouth by 41.5%. A 1998 study by the
University of Athens The National and Kapodistrian University of Athens (NKUA; el, Εθνικό και Καποδιστριακό Πανεπιστήμιο Αθηνών, ''Ethnikó ke Kapodistriakó Panepistímio Athinón''), usually referred to simply as the Univers ...
found that mastic oil has antibacterial and antifungal properties. Another 1998 University of Nottingham study claims that mastic can heal
peptic ulcers Peptic ulcer disease (PUD) is a break in the inner lining of the stomach, the first part of the small intestine, or sometimes the lower esophagus. An ulcer in the stomach is called a gastric ulcer, while one in the first part of the intestines i ...
by killing ''
Helicobacter pylori ''Helicobacter pylori'', previously known as ''Campylobacter pylori'', is a gram-negative, microaerophilic, spiral (helical) bacterium usually found in the stomach. Its helical shape (from which the genus name, helicobacter, derives) is though ...
'', which causes peptic ulcers,
gastritis Gastritis is inflammation of the lining of the stomach. It may occur as a short episode or may be of a long duration. There may be no symptoms but, when symptoms are present, the most common is upper abdominal pain (see dyspepsia). Other possi ...
, and
duodenitis Duodenitis is inflammation of the duodenum. It may persist acutely or chronically. Symptoms Known symptoms of duodenitis include: * Abdominal pain * vomiting * nausea * discomfort in stomach Causes Known causes of duodenitis include: * ''Hel ...
. Some ''in vivo'' studies have shown that mastic gum has no effect on ''H. pylori'' when taken for short periods of time. However, a recent and more extensive study showed that mastic gum reduced ''H. pylori'' populations after an insoluble and sticky polymer (poly-β-myrcene) constituent of mastic gum was removed, and if taken for a longer period of time.


Miscellanea

Apart from its medicinal properties and cosmetic and culinary uses, mastic gum is also used in the production of high-grade
varnish Varnish is a clear transparent hard protective coating or film. It is not a stain. It usually has a yellowish shade from the manufacturing process and materials used, but it may also be pigmented as desired, and is sold commercially in various ...
. The mastic tree has been introduced into
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
as an
ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...
, where it is very prized and fully
naturalized Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...
. The trees are grown mainly in suburban areas in semiarid zones, and remain undamaged, although the summer rainfall is contrary to its original
Mediterranean climate A Mediterranean climate (also called a dry summer temperate climate ''Cs'') is a temperate climate sub-type, generally characterized by warm, dry summers and mild, fairly wet winters; these weather conditions are typically experienced in the ...
. A related species, ''P. saportae'', has been shown by DNA analysisTingshuang Yi, Jun Wen, Avi Golan-Goldhirsh, and Dan E. Parfitt. Phylogenetics and reticulate evolution in PISTACIA ( Anacardiaceae). American Journal of Botany 95(2): 241–251. 2008. http://www.amjbot.org/content/95/2/241.full.pdf to be a hybrid between maternal ''P. lentiscus'' and paternal ''P. terebinthus'' (terebinth or turpentine). The hybrid has imparipinnate leaves, with leaflets semipersistent, subsessile terminal, and sometimes reduced. Usually, ''P. terebinthus'' and'' P. lentiscus'' occupy different biotopes and barely overlap: Mastic appears at lower elevations and near the sea, while the ''P. terebinthus'' most frequently inhabits inland and mountainous areas such as the
Iberian System Iberian refers to Iberia. Most commonly Iberian refers to: *Someone or something originating in the Iberian Peninsula, namely from Spain, Portugal, Gibraltar and Andorra. The term ''Iberian'' is also used to refer to anything pertaining to the fo ...
. "Dufte-Zeichen" (Scents-signs), the fourth scene from ''
Sonntag aus Licht (Sunday from Light) is an opera by Karlheinz Stockhausen in five scenes and a farewell, to a libretto written and compiled by the composer. It is the last-composed of seven operas that comprise the cycle '' Licht'' (Light). Its stage premiere in 2 ...
'' by
Karlheinz Stockhausen Karlheinz Stockhausen (; 22 August 1928 – 5 December 2007) was a German composer, widely acknowledged by critics as one of the most important but also controversial composers of the 20th and early 21st centuries. He is known for his groun ...
, is centred around seven scents, each one associated with one day of the week. "Mastix" is assigned to Wednesday and comes third.


See also

* False mastic *
Greek cuisine Greek cuisine (Greek: Ελληνική Κουζίνα) is the cuisine of Greece and the Greek diaspora. In common with many other cuisines of the Mediterranean, it is founded on the triad of wheat, olive oil, and wine. It uses vegetables, oli ...
*
Greek food products Greece produces many food products. Olive oil Greece is the world's fifth ranked producer of olive oil, producing more than 1,079,000 tons of olive oil annually, more than 75% of that extra virgin. Greek olive oil is exported throughout the worl ...
*
Mastic (plant resin) Mastic ( el, Μαστίχα) is a resin obtained from the mastic tree (''Pistacia lentiscus''). It is also known as tears of Chios, being traditionally produced on the island Chios, and, like other natural resins, is produced in "tears" or dropl ...
*
Mastichochoria Mastichochoria ( el, Μαστιχοχώρια, , lit. "the mastic villages" in English) is a former municipality on the island of Chios (Χίος), North Aegean, Greece. Since the 2011 local government reform it is part of the municipality Chios, ...
*
Turkish cuisine Turkish cuisine () is the cuisine of Turkey and the Turkish diaspora. It is largely the heritage of Ottoman cuisine, which can be described as a fusion and refinement of Mediterranean, Balkan, Middle Eastern, Central Asian and Eastern Eur ...


References


Further reading

* {{Taxonbar, from=Q159812 lentiscus Flora of North Africa Flora of Western Asia Trees of Europe Greek cuisine Resins Spices Chios Trees of Mediterranean climate Garden plants of Europe Garden plants of Africa Garden plants of Asia Drought-tolerant trees Ornamental trees Plants described in 1753 Taxa named by Carl Linnaeus Flora of the Mediterranean Basin