The Lebanon viper (''Montivipera bornmuelleri''), also known as Bornmueller's viper, Mallow D,

Ludwig D Ludwig may refer to: People and fictional characters * Ludwig (given name), including a list of people and fictional characters * Ludwig (surname), including a list of people * Ludwig Ahgren, or simply Ludwig, American YouTube live streamer and co ...

, Nilson G (2003). ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. 359 pp. . is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

venomous snake Venomous snakes are Species (biology), species of the Suborder (biology), suborder Snake, Serpentes that are capable of producing Snake venom, venom, which they use for killing prey, for defense, and to assist with digestion of their prey. The v ...

in the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

Viperinae The Viperinae, or viperines, are a subfamily of vipers endemic to Europe, Asia and Africa. They are distinguished by their lack of the heat-sensing pit organs that characterize their sister group, the subfamily Crotalinae. Currently, 13 genera a ...

of the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Viperidae The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...

. The species is native to

Western Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of Asia, as defined by some academics, UN bodies and other institutions. It is almost entirely a part of the Middle East, and includes Ana ...

. www.reptile-database.org. There are no recognized

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

Etymology

The specific name, ''bornmuelleri'', is in honor of

German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...

botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...

Joseph Friedrich Nicolaus Bornmüller.

Description

''M. bornmuelleri'' grows to a maximum total length (including tail) of about , but usually much less. Males tend to be larger than females in some populations. In specimens from

Mount Lebanon Mount Lebanon ( ar, جَبَل لُبْنَان, ''jabal lubnān'', ; syr, ܛܘܪ ܠܒ݂ܢܢ, ', , ''ṭūr lewnōn'' french: Mont Liban) is a mountain range in Lebanon. It averages above in elevation, with its peak at . Geography The Mount Le ...

, the maximum total lengths were for females and for males. The tail accounts for about 7–10% of the total length.

Geographic range and habitat

''M. bornmuelleri'' is found in high mountain areas in

Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...

Lebanon Lebanon ( , ar, لُبْنَان, translit=lubnān, ), officially the Republic of Lebanon () or the Lebanese Republic, is a country in Western Asia. It is located between Syria to the north and east and Israel to the south, while Cyprus li ...

, and

Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...

. Its preferred natural

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s are cedar

forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...

shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It m ...

, and alpine

grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...

Reproduction

''M. bornmuelleri'' is

viviparous Among animals, viviparity is development of the embryo inside the body of the parent. This is opposed to oviparity which is a reproductive mode in which females lay developing eggs that complete their development and hatch externally from the m ...

Taxonomy

The original

syntype In biological nomenclature, a syntype is any one of two or more biological types that is listed in a description of a taxon where no holotype was designated. Precise definitions of this and related terms for types have been established as part of ...

s of ''M. bornmuelleri'' were collected in Lebanon at 1800 m (5,900 ft) and in the Bolkar mountains of western

Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...

at 2200 m (7,200 ft) as described by

Franz Werner Franz Josef Maria Werner (15 August 1867 in Vienna – 28 February 1939 in Vienna) was an Austrian zoologist and explorer. Specializing as a herpetologist and entomologist, Werner described numerous species and other taxa of frogs, snakes, inse ...

in 1898.Nilson, Göran; Andrén, Claes (1985). "Systematics of the ''Vipera xanthina'' Complex (Reptilia: Viperidae). III. Taxonomic Status of the Bulgar Dagh Viper in South Turkey". ''Journal of Herpetology'' 19 (2): 276–283. (p. 276.) In 1922, Werner restricted the type locality to Lebanon in his designation of his specimen as a

lectotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...

, and in 1938 separated out the southern varieties as a separate species (''

Daboia palaestinae ''Daboia palaestinae'', also known as the Palestine viper,Mallow D, Ludwig D, Nilson G. 2003. ''True Vipers: Natural History and Toxinology of Old World Vipers.'' Malabar, Florida: Krieger Publishing Company. 359 pp. .Mehrtens JM. 1987. ''Living ...

''). In 1967

Mertens __NOTOC__ Mertens () is a surname of Flemish Origin, meaning "son of Merten" (Martin). It is the fifth most common name in Belgium with 18,518 people in 2008. Geographical distribution As of 2014, 43.4% of all known bearers of the surname ''Merte ...

raised the name ''bornmuelleri'' to valid species rank for the Lebanese populations, thus leaving the name ''xanthina'' for all Turkish populations, which arrangement was accepted by Baran in 1976, Baran İ (1976). "''Tiirkiye yilanlarinin taksonomik revizyonu ve cografi dagilislari'' ". ''TBTAK Yayinlari, Ankara, T.B.A.G.'' Series 9, (309): 177. (in Turkish). and agreed with by Nilson and Andrén in their 1985 paper.

Conservation status

The species ''M. bornmuelleri'' is classified as Endangered (EN) according to the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

Red List of Threatened Species with the following criteria: B1ab(iii) (v3.1, 2001). This indicates that it is facing a high risk of extinction in the wild because the extent of its occurrence within its geographic range is estimated to be less than 5,000 km² (1,931 sq mi), because its populations are severely fragmented or known to exist at no more than 5 locations, and because a continued decline is observed, inferred or projected in the area, extent and/or quality of its habitat.