Lansium Domesticum
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''Lansium parasiticum'', commonly known as langsat (), lanzones (), or longkong in English; duku in
Indonesian Indonesian is anything of, from, or related to Indonesia, an archipelagic country in Southeast Asia. It may refer to: * Indonesians, citizens of Indonesia ** Native Indonesians, diverse groups of local inhabitants of the archipelago ** Indonesian ...
or dokong in
Terengganu Malay Terengganu Malay ( ms, Bahasa Melayu Terengganu; Terengganu Malay: ) is a Malayic language spoken in the Malaysian state of Terengganu all the way southward to coastal Pahang and northeast Johor. It is the native language of Terengganu Malays an ...
, is a species of tree in the
Mahogany Mahogany is a straight-grained, reddish-brown timber of three tropical hardwood species of the genus ''Swietenia'', indigenous to the AmericasBridgewater, Samuel (2012). ''A Natural History of Belize: Inside the Maya Forest''. Austin: Unive ...
family with commercially cultivated edible fruits. The species is native to Southeast Asia.


Description

The tree is average sized, reaching in height and in diameter. 30 years old trees grown from seed and planted at 8 × 8 meter spacing can have a height of 10 meters and diameter of 25 cm. The trunk grows in an irregular manner, with its buttress roots showing above ground. The tree's bark is a greyish colour, with light and dark spots. Its resin is thick and milk coloured. The
pinnately compound Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
leaves are odd numbered, with thin hair, and 6 to 9 buds at intervals. The buds are long and elliptical, approximately by in size. The upper edge shines, and the leaves themselves have pointed bases and tips. The stems of the buds measure . The flowers are located in inflorescences that grow and hang from large branches or the trunk; the bunches may number up to 5 in one place. They are often branched at their base, measure in size, and have short fur. The flowers are small, with short stems, and have two
genders Gender is the range of characteristics pertaining to femininity and masculinity and differentiating between them. Depending on the context, this may include sex-based social structures (i.e. gender roles) and gender identity. Most cultures u ...
. The sheathe is shaped like a five lobed cup and is coloured a greenish-yellow. The corona is egg-shaped and hard, measuring by . There is one
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
, measuring in length. The top of the stamen is round. The
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
is short and thick. The fruit can be elliptical, ovoid or round, measuring by in size. Fruits look much like small potatoes and are borne in clusters similar to grapes. The larger fruits are on the variety known as duku. It is covered by thin, yellow hair giving a slightly fuzzy aspect. The skin thickness varies with the varieties, from to approximately . With the skin removed, the fruit resembles peeled
garlic Garlic (''Allium sativum'') is a species of bulbous flowering plant in the genus ''Allium''. Its close relatives include the onion, shallot, leek, chive, Allium fistulosum, Welsh onion and Allium chinense, Chinese onion. It is native to South A ...
in appearance, with half a dozen white, translucent lobes, some of which contain a flat, bitter tasting seed. The seeds are covered with a thick, clear-white
aril An aril (pronounced ), also called an arillus, is a specialized outgrowth from a seed that partly or completely covers the seed. An arillode or false aril is sometimes distinguished: whereas an aril grows from the attachment point of the see ...
that tastes sweet and sour. The taste has been likened to a combination of grape and grapefruit and is considered excellent by most. The sweet juicy flesh contains
sucrose Sucrose, a disaccharide, is a sugar composed of glucose and fructose subunits. It is produced naturally in plants and is the main constituent of white sugar. It has the molecular formula . For human consumption, sucrose is extracted and refined ...
,
fructose Fructose, or fruit sugar, is a Ketose, ketonic monosaccharide, simple sugar found in many plants, where it is often bonded to glucose to form the disaccharide sucrose. It is one of the three dietary monosaccharides, along with glucose and galacto ...
, and glucose. For consumption, cultivars with small or undeveloped seeds and thick aril are preferred.


Cultivars

There are numerous cultivars of ''L. parasiticum''. Overall, there are two main groups of cultivars, those named ''duku'' and those named ''langsat''. There are also mixed duku-langsat varieties. Those called duku generally have a large crown, thick with bright green leaves, with short bunches of few fruit. The individual fruit are large, generally round, and have somewhat thick skin that does not release sap when cooked. The seeds are small, with thick flesh, a sweet scent, and a sweet or sour taste. Meanwhile, the variant commonly known as langsat generally has thinner trees, with a less dense crown consisting of dark green leaves and stiff branches. The bunches are longer, and each bunch holds between 15 and 25 large, egg-shaped fruit. The skin is thin and releases a white sap when cooked. The flesh is watery and tastes sweet and sour. ''Lansium domesticum'' var. ''aquaeum'' is distinguished by its hairy leaves, as well as the tightly packed dark yellow fruit on its bunches. The fruit tends to be small, with thin skin and little sap; the skin is difficult to remove. To be eaten, the fruit is bitten and the flesh sucked through the hole created, or rubbed until the skin breaks and the seeds are retrieved. In Indonesia the fruit has several names, including ''kokosan'', ''pisitan'', ''pijetan'', and ''bijitan''. The seeds are relatively large, with thin, sour flesh.


Reproduction

The seeds of ''L. parasiticum'' are polyembryonic, the multiple embryos resulting from
apomixis In botany, apomixis is asexual reproduction without fertilization. Its etymology is Greek for "away from" + "mixing". This definition notably does not mention meiosis. Thus "normal asexual reproduction" of plants, such as propagation from cuttin ...
. ''Lansium parasiticum'' is traditionally reproduced by spreading seedlings, either cultivated or collected from below the tree. It has been said that new seedlings require 20 to 25 years to bear fruit, with the possibility of the quality being inferior. However other sources quote 12 years to first production from seed and no variations. Production often varies from year to year, and depends to some extent on having a dry period to induce flowering. One example of ten trees in Costa Rica about twenty-five years old produced during five years the following weights of salable fruits: 2008: 50 kilos, 2009: 2000 kilos, 2010: 1000 kilos, 2011: 100 kilos, 2012: 1500 kilos. Experiments in the Philippines with grafting where two trees are planted close to each other and then grafted when one to two meters tall to leave twin root systems on a single main trunk have resulted in earlier and less erratic fruit production. Another common method is by
air layering Layering has evolved as a common means of vegetative propagation of numerous species in natural environments. Layering is also utilized by horticulturists to propagate desirable plants. Natural layering typically occurs when a branch touches ...
. Although the process requires up to several months, the new rooted tree produced is itself ready to bear fruit within two years. Trees cultivated with this method have a high death rate, and the growths are less resilient. The third common way to reproduce ''L. parasiticum'' is with
grafting Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the rootstock. The succ ...
. This results in the new trees having the same genetic characteristics as their parent, and being ready to bear fruit within 5 to 6 years. The offspring are relatively stronger than transplanted shoots.


Ecology

''Lansium parasiticum'' grows well in mixed agroforests. The plant, especially the ''duku'' variant, prefers damp, shaded areas. It can be grown in the same agroforest as
durian The durian (, ) is the edible fruit of several tree species belonging to the genus ''Durio''. There are 30 recognised ''Durio'' species, at least nine of which produce edible fruit. ''Durio zibethinus'', native to Borneo and Sumatra, is the onl ...
,
petai ''Parkia speciosa'', the bitter bean, twisted cluster bean or stink bean, is a plant of the genus ''Parkia'' in the family Fabaceae. It bears long, flat edible beans with bright green seeds the size and shape of plump almonds which have a ra ...
, and
jengkol ''Archidendron pauciflorum'', commonly known as djenkol, jengkol or jering is a species of flowering tree in the pea family, Fabaceae. It is native to Southeast Asia, where the seeds are a popular dish.Lim, T. K. "Archidendron jiringa." Edible Me ...
, as well as wood-producing trees. ''Lansium parasiticum'' is grown from low grounds up to heights of above sea level, in areas with an average rainfall of a year. The plant can grow and blossom in
latosol Latosols, also known as tropical red earth, are soils found under tropical rainforests which have a relatively high content of iron and aluminium oxides. They are typically classified as oxisols (USDA soil taxonomy) or ferralsols (World Reference B ...
, yellow podzol, and alluvium. The plant prefers slightly acidic soil with good drainage and rich in mulch. The ''langsat'' variant is hardier, and can weather dry seasons with a little shade and water. The plant cannot handle floods. ''Lansium parasiticum'' generally bears fruit once a year. This period can vary between areas, but blooming is generally after the beginning of the rainy season and fruit production some four months later.


Distribution

''Lansium parasiticum'' is native to the Malesian
phytochorion A phytochorion, in phytogeography, is a geographic area with a relatively uniform composition of plant species. Adjacent phytochoria do not usually have a sharp boundary, but rather a soft one, a transitional area in which many species from both re ...
of Southeast Asia, from
Peninsular Thailand Southern Thailand, Southern Siam or Tambralinga is a southernmost cultural region of Thailand, separated from Central Thailand region by the Kra Isthmus. Geography Southern Thailand is on the Malay Peninsula, with an area of around , bounded ...
and Peninsular Malaysia, to Java,
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
, Borneo, the Philippines ( Luzon, Camiguin, Basilan, and Mindanao),
Sulawesi Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu Ar ...
, the Moluccas, and Western New Guinea. It has also been introduced to
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
, Cambodia,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
, Micronesia, Hawaii,
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
, India, the Seychelles, Trinidad and Tobago, and
Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the north ...
, among others. The tree is cultivated commercially in Thailand ( th, ลางสาด, langsat), Cambodia, Vietnam, India, Malaysia and the Philippines ( tl, lansones; ceb, buwahan, lansones; Philippine Spanish: ''lanzones''). It grows well in the wetter areas (120 inches/3 meters or more annual rainfall) of Costa Rica, where it is still very rare, having been introduced decades ago by the United Fruit Company. A major hindrance to its acceptance seems to be that it is very slow in bearing, said to take 12 years or more from seed. However, air layering from mature trees, as well as grafting, are said to work well and produce much faster. The name Duku is reserved for the larger-sized varieties of Langsat, near the size of golf balls, claimed sweeter and with less sap in the peel. A variety called Dokong exported to mainland Malaysia from Thailand (this variety is called 'Longkang' th, ลองกอง in Thailand) grows tighter in the clusters, giving it a faceted shape, and is preferred by many over the standard Langsat.


Uses

''Lansium parasiticum'' is cultivated mainly for its fruit, which can be eaten raw. The fruit can also be bottled in syrup. The wood is hard, thick, heavy, and resilient, allowing it to be used in the construction of rural houses. Some parts of the plant are used in making traditional medicine. The bitter seeds can be pounded and mixed with water to make a deworming and ulcer medication. The bark is used to treat dysentery and malaria; the powdered bark can also be used to treat scorpion stings. The fruit's skin is used to treat diarrhea, and in the Philippines the dried skin is burned as a mosquito repellent. The skin, especially of the ''langsat'' variety, can be dried and burned as incense. The greatest producers of ''Lansium parasiticum'' are Indonesia, Thailand, the Philippines and Malaysia. The production is mostly for internal consumption, although some is exported to Singapore, Hong Kong and Kuwait.


See also

*'' Dysoxylum parasiticum'' * Lychee * Longan


References


External links

* *
''Lansium domesticum''
at Tree Functional Attributes and Ecological Database (World Agro Forestry) {{Taxonbar, from=Q860731 Meliaceae Tropical fruit Fruits originating in Asia Trees of Indo-China Trees of Malesia Fruit trees