Lake Chub
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The lake chub (''Couesius plumbeus'') is a
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
cyprinid Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest verte ...
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
found in
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
and in parts of the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
. Of all
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
n
minnow Minnow is the common name for a number of species of small freshwater fish, belonging to several genera of the families Cyprinidae and Leuciscidae. They are also known in Ireland as pinkeens. Smaller fish in the subfamily Leusciscidae are co ...
s, it is the one with the northernmost distribution. Its
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
, ''Couesius'' is considered
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
today. The genus was named after
Elliott Coues Elliott Ladd Coues (; September 9, 1842 – December 25, 1899) was an American army surgeon, historian, ornithologist, and author. He led surveys of the Arizona Territory, and later as secretary of the United States Geological and Geographic ...
, who collected the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen.


Description

The body is
fusiform Fusiform means having a spindle-like shape that is wide in the middle and tapers at both ends. It is similar to the lemon-shape, but often implies a focal broadening of a structure that continues from one or both ends, such as an aneurysm on a b ...
and somewhat elongate. It is usually 100 to 175 mm (4 to 7 in) long, where the maximum length is approximately 225 mm (9 in). The back is olive-brown or
dark brown Brown is a color. It can be considered a composite color, but it is mainly a darker shade of orange. In the CMYK color model used in printing or painting, brown is usually made by combining the colors orange and black. In the RGB color model use ...
, and the sides are leaden silver, hence the word ''plumbeus'', referring to lead, in the scientific name of this fish. The snout is blunt and projects slightly beyond the upper lip. The corners of the mouth each bear a small barbel. The scales are small but well visible, and some may be black and form isolated dark spots on the lower sides. The dorsal, pelvic and anal fins each have 8 rays. The pectoral fins are broad and have 13-18, but more usually 15-16, rays. The fish has no
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
, adipose fin, or spines. The mouth is relatively small and the tail is forked. Breeding males can develop patches of bright orange or red at the base of the pectoral fins and sometimes near the mouth, and small nuptial tubercles on the top of the head, dorsal surface of pectoral rays, and on the belly near the base of the pectorals. The juveniles are similar to the adults, but have less obvious mottling or stripe. The females are typically larger than the males.


Distribution

The lake chub is generally found throughout
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
up to the
Arctic Circle The Arctic Circle is one of the two polar circles, and the most northerly of the five major circles of latitude as shown on maps of Earth. Its southern equivalent is the Antarctic Circle. The Arctic Circle marks the southernmost latitude at w ...
. Some scattered populations are also present in the
north North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating Direction (geometry), direction or geography. Etymology T ...
ern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
, more precisely in
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
,
Michigan Michigan () is a state in the Great Lakes region of the upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the 10th-largest state by population, the 11th-largest by area, and the ...
,
Iowa Iowa () is a state in the Midwestern region of the United States, bordered by the Mississippi River to the east and the Missouri River and Big Sioux River to the west. It is bordered by six states: Wisconsin to the northeast, Illinois to the ...
,
Wisconsin Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake M ...
,
Minnesota Minnesota () is a state in the upper midwestern region of the United States. It is the 12th largest U.S. state in area and the 22nd most populous, with over 5.75 million residents. Minnesota is home to western prairies, now given over to ...
, the Dakotas, Montana,
Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...
,
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
,
Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...
,
Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyom ...
and
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
. The lake chub is also the only
minnow Minnow is the common name for a number of species of small freshwater fish, belonging to several genera of the families Cyprinidae and Leuciscidae. They are also known in Ireland as pinkeens. Smaller fish in the subfamily Leusciscidae are co ...
known to live in
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
. Its range also includes
Lake Michigan Lake Michigan is one of the five Great Lakes of North America. It is the second-largest of the Great Lakes by volume () and the third-largest by surface area (), after Lake Superior and Lake Huron. To the east, its basin is conjoined with that o ...
in
Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolita ...
and the
Platte River The Platte River () is a major river in the State of Nebraska. It is about long; measured to its farthest source via its tributary, the North Platte River, it flows for over . The Platte River is a tributary of the Missouri River, which itself ...
system in Wyoming. There are
relict A relict is a surviving remnant of a natural phenomenon. Biology A relict (or relic) is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas. Geology and geomorphology In geology, a r ...
populations of the lake chub in the upper Missouri River
drainage Drainage is the natural or artificial removal of a surface's water and sub-surface water from an area with excess of water. The internal drainage of most agricultural soils is good enough to prevent severe waterlogging (anaerobic conditio ...
.


Habitat

As its
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
implies, the lake chub is most commonly found in cold-water
lake A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much large ...
s with clean
gravel Gravel is a loose aggregation of rock fragments. Gravel occurs naturally throughout the world as a result of sedimentary and erosive geologic processes; it is also produced in large quantities commercially as crushed stone. Gravel is classifi ...
, and it can also live in cold-water
river A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of wate ...
s and
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a stream ...
s. It often lives in shallows, but during mid-
summer Summer is the hottest of the four temperate seasons, occurring after spring and before autumn. At or centred on the summer solstice, the earliest sunrise and latest sunset occurs, daylight hours are longest and dark hours are shortest, wit ...
, it may move to the deeper parts of a
lake A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much large ...
to avoid the warmer waters of the
lake A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much large ...
shore A shore or a shoreline is the fringe of land at the edge of a large body of water, such as an ocean, sea, or lake. In physical oceanography, a shore is the wider fringe that is geologically modified by the action of the body of water past a ...
.


Diet

The diet of the lake chub is varied:
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
, insects, aquatic insect larvae, and algae. The largest individuals can capture small fishes. The lake chub itself can be eaten by large predatory fishes and is therefore suitable as bait for fishing.


Growth

In and around Catamaran Brook, New Brunswick, Canada, lake chubs over 5 cm long (2 inches) were found to grow by about 0.8 cm (0.3 inch) a month in summer.


Reproduction

During spawning, male lake chub will pursue females. Spawning-ready males will charge a female which causes her to swim upward, sometimes breaking the water surface. The male swims next to the female, and if multiple males are present, they swim next to one another in order to compete for the female. During this process, the male's mouth will open and close rapidly until the female swims to a rock and the remaining male moves against her until her eggs are released. Lake chubs normally undergo spawning migrations in early summer. Temperature plays a role in triggering migration onset, as migrations are delayed on colder years. Migrating chubs leave their lakes and rivers to ascend tributary streams, in which they mate (by day or night) and release their eggs over gravel or rocks. There is no parental care. During migration the fish move mostly during dusk and at night.


Behavior

In the laboratory, lake chubs have expressed free-running
circadian rhythm A circadian rhythm (), or circadian cycle, is a natural, internal process that regulates the sleep–wake cycle and repeats roughly every 24 hours. It can refer to any process that originates within an organism (i.e., Endogeny (biology), endogeno ...
s that are among the most precise of the few fish species studied to date. These fish are usually diurnal in the laboratory, but in the wild they can be diurnal, crepuscular, or nocturnal. The lake chub has large optic lobes in its brain and is therefore presumed to be a good sight feeder. It can, however, feed at night also. Comparisons with other freshwater fishes such as
stickleback The sticklebacks are a family of ray-finned fishes, the Gasterosteidae which have a Holarctic distribution in fresh, brackish and marine waters. They were thought to be related to the pipefish and seahorses but are now thought to be more closel ...
,
northern pike The northern pike (''Esox lucius'') is a species of carnivorous fish of the genus '' Esox'' (the pikes). They are typical of brackish and fresh waters of the Northern Hemisphere (''i.e.'' holarctic in distribution). They are known simply as a ...
,
sculpin A sculpin is a type of fish that belongs to the superfamily Cottoidea in the order Scorpaeniformes.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology suggest transitions in station-holding demand a ...
, and
burbot The burbot (''Lota lota'') is the only gadiform (cod-like) freshwater fish Freshwater fish are those that spend some or all of their lives in fresh water, such as rivers and lakes, with a salinity of less than 1.05%. These environments di ...
have revealed that it has a superior hearing capacity.Mann, D.A., Cott, P.A., Hanna, B.W., and Popper, A.N., 2007, Hearing in eight species of northern Canadian freshwater fishes, Journal of Fish Biology 70: 109-120.


Common name

The lake chub is sometimes called by other names such as northern chub, lake northern chub, chub minnow, plumbeus minnow, or bottlefish. The species is also found in Quebec and its French name is "mené de lac". The lake chub must not be confused with various
cisco Cisco Systems, Inc., commonly known as Cisco, is an American-based multinational digital communications technology conglomerate corporation headquartered in San Jose, California. Cisco develops, manufactures, and sells networking hardware, ...
species of the genus ''
Coregonus ''Coregonus'' is a diverse genus of fish in the salmon family (Salmonidae). The ''Coregonus'' species are known as whitefishes. The genus contains at least 68 described extant taxa, but the true number of species is a matter of debate. The type ...
'' living in
Lake Michigan Lake Michigan is one of the five Great Lakes of North America. It is the second-largest of the Great Lakes by volume () and the third-largest by surface area (), after Lake Superior and Lake Huron. To the east, its basin is conjoined with that o ...
, which are often called Michigan Lake chubs.


Identification


Subspecies

Three distinct subspecies of lake chub have been identified: ''C. p plumbeus'', ''C. p greeni'', and one formerly known as ''C. p dissimilis''. A name is no longer available for the ''C. p dissimilis''.


References

*


External links


FishBase valid species list
{{Taxonbar, from=Q2540189
lake chub The lake chub (''Couesius plumbeus'') is a freshwater cyprinid fish found in Canada and in parts of the United States. Of all North American minnows, it is the one with the northernmost distribution. Its genus, ''Couesius'' is considered mono ...
lake chub The lake chub (''Couesius plumbeus'') is a freshwater cyprinid fish found in Canada and in parts of the United States. Of all North American minnows, it is the one with the northernmost distribution. Its genus, ''Couesius'' is considered mono ...
Lake fish of North America Freshwater fish of the United States Fish of Canada Fish of the Great Lakes
lake chub The lake chub (''Couesius plumbeus'') is a freshwater cyprinid fish found in Canada and in parts of the United States. Of all North American minnows, it is the one with the northernmost distribution. Its genus, ''Couesius'' is considered mono ...