Labrisomids are small blennioids (blennies), percomorph marine

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

belonging to the family Labrisomidae. Found mostly in the tropical

Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...

and

Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

, the family contains about 110 species in 15 genera. Stockier than the average blenny, labrisomids are elongated nonetheless; their dorsal fin spines outnumber soft

rays Ray may refer to: Fish * Ray (fish), any cartilaginous fish of the superorder Batoidea * Ray (fish fin anatomy), a bony or horny spine on a fin Science and mathematics * Ray (geometry), half of a line proceeding from an initial point * Ray (gra ...

(which may be absent altogether), and their

pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two en ...

s are long and slender. Like many other blennies, labrisomids have whisker-like structures called

cirri Giovanni Battista Cirri (1 October 1724 – 11 June 1808) was an Italian cellist and composer in the 18th century. Biography Cirri was born in Forlì in the Emilia-Romagna Region of Italy. He had his first musical training with his brother ...

on their heads and

napes ''N''-Acylphosphatidylethanolamines (NAPEs) are hormones released by the small intestine into the bloodstream when it processes fat. NAPEs travel to the hypothalamus in the brain and suppress appetite. This mechanism could be relevant for treating ...

. Scales may be

cycloid In geometry, a cycloid is the curve traced by a point on a circle as it rolls along a straight line without slipping. A cycloid is a specific form of trochoid and is an example of a roulette, a curve generated by a curve rolling on another curve ...

or absent in labrisomids; many species are brightly coloured. The

hairy blenny ''Labrisomus nuchipinnis'', the hairy blenny, is a species of labrisomid blenny native to the Atlantic Ocean from the coast of the Americas to the African coast. This species prefers areas that give them crevices and holes to shelter in such as ...

(''Labrisomus nuchipinnis'') is the largest species at 23 cm in length; most are far smaller. Generally staying within shallow coastal regions to depths around 10 m, labrisomids are

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

fish spending most of their time on or near the bottom. Both sandy and rocky substrates are frequented, sometimes at

reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic processes— deposition of sand, wave erosion planing down rock out ...

s or amongst beds of

seagrass Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families (Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the orde ...

. Labrisomids are shy fish and will retreat into crevices if threatened.

Crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...

brittle star Brittle stars, serpent stars, or ophiuroids (; ; referring to the serpent-like arms of the brittle star) are echinoderms in the class Ophiuroidea, closely related to starfish. They crawl across the sea floor using their flexible arms for locomo ...

s, and

sea urchin Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) of ...

s make up much of the labrisomid diet. Two genera of labrisomid are noted for their

ovoviviparity Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...

; ''

Xenomedea ''Xenomedea rhodopyga'', common name the redrump blenny, is a species of labrisomid blenny endemic to the Gulf of California. It inhabits weed-covered rocky reefs and tide pools A tide pool or rock pool is a shallow pool of seawater tha ...

'' and ''

Starksia ''Starksia'' is a genus of labrisomid blennies native to the western Atlantic Ocean and the eastern Pacific Ocean. Their typical length is SL. The generic name honours the American ichthyologist Edwin Chapin Starks (1867-1932) of Stanford Un ...

'' both retain eggs within their

oviduct The oviduct in mammals, is the passageway from an ovary. In human females this is more usually known as the Fallopian tube or uterine tube. The eggs travel along the oviduct. These eggs will either be fertilized by spermatozoa to become a zygote, o ...

s, where they develop in safety. However, only ''Starksia'' species possess gonopodia (modified anal fins used as a copulatory organ). Some workers have found that the Labrisomidae is

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

Genera

The following genera are currently included in this family: * ''

Alloclinus ''Alloclinus holderi'', the island kelpfish is a subtropical species of labrisomid blenny native to the eastern Pacific Ocean from Santa Cruz Island, California, to Baja California. This species inhabits rocky areas and can be found down to abo ...

'' C.L. Hubbs, 1927 * ''

Auchenionchus ''Auchenionchus'' is a genus of labrisomid blennies endemic to the Pacific waters off of Chile, Species ''Auchenionchus'' contains these currently recognized species: * '' Auchenionchus crinitus'' ( Jenyns, 1841) * '' Auchenionchus microcirrhis' ...

Gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...

, 1860 * ''

Brockius ''Brockius'' is a genus of labrisomid blennies from the waters of the eastern Pacific and the western Atlantic where they are associated with reefs and seaweed-covered rocks. Species The species in the genus are: *'' Brockius albigenys'' ( Beebe ...

'' Clark L. Hubbs, 1953 * ''

Calliclinus ''Calliclinus'' is a genus of labrisomid blennies native to the Pacific and Atlantic coasts of southern South America. Species There are currently two recognized species in this genus: * '' Calliclinus geniguttatus'' (Valenciennes, 1836) * '' Ca ...

'' Gill, 1860 * ''

Cottoclinus ''Cottoclinus canops'' is a species of labrisomid blenny endemic to the Galápagos Islands. Males of this species can reach a length of SL while females can reach a length of . It is the only known member of its genus Genus ( plural gener ...

'' McCosker,

Stephens Stephens is a surname. It is a patronymic and is recorded in England from 1086. Notable people with the surname include: *Alexander H. Stephens (1812–1883), Vice President of the Confederate States of America *Alison Stephens (1970–2010), Brit ...

Rosenblatt Rosenblatt is a surname of German and Jewish origin, meaning "rose leaf". People with this surname include: * Albert Rosenblatt (born 1936), New York Court of Appeals judge * Dana Rosenblatt, known as "Dangerous" (born 1972), American boxer * Elie ...

, 2003 * '' Cryptotrema''

Gilbert Gilbert may refer to: People and fictional characters * Gilbert (given name), including a list of people and fictional characters *Gilbert (surname), including a list of people Places Australia * Gilbert River (Queensland) * Gilbert River (South ...

, 1890 * '' Dialommus'' Gilbert, 1891 * '' Exerpes''

Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...

& Evermann, 1896 * ''

Gobioclinus ''Gobioclinus'' is a genus of labrisomid blennies from the coasts of the western Atlantic and eastern Pacific Oceans off the Americas. Species The following species are classified within the genus ''Gobioclinus'': *'' Gobioclinus bucciferus'' ( ...

'' Gill, 1860 * '' Haptoclinus'' Böhlke &

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, 1974 * ''

Labrisomus ''Labrisomus'' is a genus of labrisomid blennies native to the western Atlantic ocean and the eastern Pacific Ocean. Species There are currently 11 recognized species in this genus: * ''Labrisomus conditus'' I. Sazima ( fr), Carvalho-Filho, ...

'' Swainson, 1839 * ''

Malacoctenus ''Malacoctenus'' is a genus of ''labrisomid'' blennies native to the eastern Pacific Ocean and the Atlantic Ocean. Species There are currently 23 recognized species in this genus: * '' Malacoctenus africanus'' Cadenat, 1951 * '' Malacoctenus au ...

'' Gill, 1860 * '' Nemaclinus'' Böhlke &

Springer Springer or springers may refer to: Publishers * Springer Science+Business Media, aka Springer International Publishing, a worldwide publishing group founded in 1842 in Germany formerly known as Springer-Verlag. ** Springer Nature, a multinationa ...

, 1975 * '' Paraclinus'' Mocquard, 1888 * ''

'' Jordan & Evermann, 1896 * ''

'' Rosenblatt &

Taylor Taylor, Taylors or Taylor's may refer to: People * Taylor (surname) **List of people with surname Taylor * Taylor (given name), including Tayla and Taylah * Taylor sept, a branch of Scottish clan Cameron * Justice Taylor (disambiguation) Plac ...

, 1971

Timeline

ImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:-65.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:5 start:-65.5 ScaleMinor = unit:year increment:1 start:-65.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -65.5 till: -55.8 color:paleocene text:

Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...

from: -55.8 till: -33.9 color:eocene text:

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

from: -33.9 till: -23.03 color:oligocene text:

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...

from: -23.03 till: -5.332 color:miocene text:

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text:

Pleist. The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...

from: -0.0117 till: 0 color:holocene text: H. bar:eratop from: -65.5 till: -23.03 color:paleogene text:

Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...

from: -23.03 till: -2.588 color:neogene text:

Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...

from: -2.588 till: 0 color:quaternary text: Q. PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:miocene bar:NAM1 from: -11.608 till: 0 text:

PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -65.5 till: -55.8 color:paleocene text:

from: -55.8 till: -33.9 color:eocene text:

from: -33.9 till: -23.03 color:oligocene text:

from: -23.03 till: -5.332 color:miocene text:

from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text:

from: -0.0117 till: 0 color:holocene text: H. bar:era from: -65.5 till: -23.03 color:paleogene text:

from: -23.03 till: -2.588 color:neogene text:

from: -2.588 till: 0 color:quaternary text: Q.

References

{{Taxonbar, from=Q183463 Blenniiformes Taxa named by Clark Hubbs