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Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
, zoologist, taxonomist, and physician who formalised binomial nomenclature, the modern system of naming organisms. He is known as the "father of modern taxonomy". Many of his writings were in Latin; his name is rendered in Latin as and, after his 1761 ennoblement, as . Linnaeus was born in
Råshult Råshult is a village just north of Älmhult in Kronoberg County, Småland, Sweden. It is notable as the birthplace of the seminal biologist and "father of modern taxonomy", Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778 ...
, the countryside of
Småland Småland () is a historical province () in southern Sweden. Småland borders Blekinge, Scania, Halland, Västergötland, Östergötland and the island Öland in the Baltic Sea. The name Småland literally means ''Small Lands''. The Latinized fo ...
, in southern Sweden. He received most of his higher education at Uppsala University and began giving lectures in botany there in 1730. He lived abroad between 1735 and 1738, where he studied and also published the first edition of his ' in the Netherlands. He then returned to Sweden where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on several journeys through Sweden to find and classify plants and animals. In the 1750s and 1760s, he continued to collect and classify animals, plants, and minerals, while publishing several volumes. He was one of the most acclaimed scientists in Europe at the time of his death. Philosopher Jean-Jacques Rousseau sent him the message: "Tell him I know no greater man on earth." Johann Wolfgang von Goethe wrote: "With the exception of Shakespeare and Spinoza, I know no one among the no longer living who has influenced me more strongly." Swedish author August Strindberg wrote: "Linnaeus was in reality a poet who happened to become a naturalist." Linnaeus has been called ' (Prince of Botanists) and "The Pliny of the North". Broberg (2006), p. 7. He is also considered one of the founders of modern ecology. In botany and zoology, the abbreviation L. is used to indicate Linnaeus as the authority for a species' name. In older publications, the abbreviation "Linn." is found. Linnaeus's remains constitute the type specimen for the species '' Homo sapiens'' following the International Code of Zoological Nomenclature, since the sole specimen that he is known to have examined was himself.ICZN Chapter 16, Article 72.4.1.1
– "For a nominal species or subspecies established before 2000, any evidence, published or unpublished, may be taken into account to determine what specimens constitute the type series." and Article 73.1.2 – "If the nominal species-group taxon is based on a single specimen, either so stated or implied in the original publication, that specimen is the holotype fixed by monotypy (see Recommendation 73F). If the taxon was established before 2000 evidence derived from outside the work itself may be taken into account rt. 72.4.1.1to help identify the specimen."


Early life


Childhood

Linnaeus was born in the village of
Råshult Råshult is a village just north of Älmhult in Kronoberg County, Småland, Sweden. It is notable as the birthplace of the seminal biologist and "father of modern taxonomy", Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778 ...
in
Småland Småland () is a historical province () in southern Sweden. Småland borders Blekinge, Scania, Halland, Västergötland, Östergötland and the island Öland in the Baltic Sea. The name Småland literally means ''Small Lands''. The Latinized fo ...
, Sweden, on 23 May 1707. He was the first child of Nicolaus (Nils) Ingemarsson (who later adopted the family name Linnaeus) and Christina Brodersonia. His siblings were Anna Maria Linnæa, Sofia Juliana Linnæa, Samuel Linnæus (who would eventually succeed their father as rector of Stenbrohult and write a manual on
beekeeping Beekeeping (or apiculture) is the maintenance of bee colonies, commonly in man-made beehives. Honey bees in the genus '' Apis'' are the most-commonly-kept species but other honey-producing bees such as ''Melipona'' stingless bees are also kept. ...
), Stöver (1794), p. 8. Broberg (2006), p. 10. and Emerentia Linnæa. His father taught him Latin as a small child. One of a long line of peasants and priests, Nils was an amateur
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
, a Lutheran
minister Minister may refer to: * Minister (Christianity), a Christian cleric ** Minister (Catholic Church) * Minister (government), a member of government who heads a ministry (government department) ** Minister without portfolio, a member of government w ...
, and the
curate A curate () is a person who is invested with the ''care'' or ''cure'' (''cura'') ''of souls'' of a parish. In this sense, "curate" means a parish priest; but in English-speaking countries the term ''curate'' is commonly used to describe clergy w ...
of the small village of Stenbrohult in Småland. Christina was the daughter of the
rector Rector (Latin for the member of a vessel's crew who steers) may refer to: Style or title *Rector (ecclesiastical), a cleric who functions as an administrative leader in some Christian denominations *Rector (academia), a senior official in an edu ...
of Stenbrohult, Samuel Brodersonius. Fries (2011), p. 376. A year after Linnaeus's birth, his grandfather Samuel Brodersonius died, and his father Nils became the rector of Stenbrohult. The family moved into the
rectory A clergy house is the residence, or former residence, of one or more priests or ministers of religion. Residences of this type can have a variety of names, such as manse, parsonage, rectory or vicarage. Function A clergy house is typically ow ...
from the curate's house. Quammen (2007), p. 1. Even in his early years, Linnaeus seemed to have a liking for plants, flowers in particular. Whenever he was upset, he was given a flower, which immediately calmed him. Nils spent much time in his garden and often showed flowers to Linnaeus and told him their names. Soon Linnaeus was given his own patch of earth where he could grow plants. Blunt (2004), p. 15. Carl's father was the first in his ancestry to adopt a permanent surname. Before that, ancestors had used the patronymic naming system of Scandinavian countries: his father was named Ingemarsson after his father Ingemar Bengtsson. When Nils was admitted to the University of Lund, he had to take on a family name. He adopted the Latinate name Linnæus after a giant linden tree (or lime tree), ' in Swedish, that grew on the family homestead. Blunt (2004), p. 12. This name was spelled with the æ ligature. When Carl was born, he was named Carl Linnæus, with his father's family name. The son also always spelled it with the æ ligature, both in handwritten documents and in publications. Blunt (2004), p. 13. Carl's patronymic would have been Nilsson, as in Carl Nilsson Linnæus.


Early education

Linnaeus's father began teaching him basic Latin, religion, and geography at an early age. When Linnaeus was seven, Nils decided to hire a tutor for him. The parents picked Johan Telander, a son of a local yeoman. Linnaeus did not like him, writing in his autobiography that Telander "was better calculated to extinguish a child's talents than develop them". Blunt (2004), pp. 15–16. Two years after his tutoring had begun, he was sent to the Lower Grammar School at Växjö in 1717. Stöver (1794), p. 5. Linnaeus rarely studied, often going to the countryside to look for plants. At some point, his father went to visit him and, after hearing critical assessements by his preceptors, he decided to put the youth as an apprentice to some honest cobbler. He reached the last year of the Lower School when he was fifteen, which was taught by the headmaster, Daniel Lannerus, who was interested in botany. Lannerus noticed Linnaeus's interest in botany and gave him the run of his garden. He also introduced him to Johan Rothman, the state doctor of Småland and a teacher at Katedralskolan (a gymnasium) in Växjö. Also a botanist, Rothman broadened Linnaeus's interest in botany and helped him develop an interest in medicine. Blunt (2004), p. 16. Stöver (1794), pp. 5–6. By the age of 17, Linnaeus had become well acquainted with the existing botanical literature. He remarks in his journal that he "read day and night, knowing like the back of my hand, Arvidh Månsson's Rydaholm Book of Herbs, Tillandz's Flora Åboensis, Palmberg's Serta Florea Suecana, Bromelii's Chloros Gothica and Rudbeckii's Hortus Upsaliensis". Linnaeus entered the Växjö Katedralskola in 1724, where he studied mainly Greek, Hebrew, theology and
mathematics Mathematics is an area of knowledge that includes the topics of numbers, formulas and related structures, shapes and the spaces in which they are contained, and quantities and their changes. These topics are represented in modern mathematics ...
, a curriculum designed for boys preparing for the priesthood. Stöver (1794), p. 6. Blunt (2004), pp. 16–17. In the last year at the gymnasium, Linnaeus's father visited to ask the professors how his son's studies were progressing; to his dismay, most said that the boy would never become a scholar. Rothman believed otherwise, suggesting Linnaeus could have a future in medicine. The doctor offered to have Linnaeus live with his family in Växjö and to teach him physiology and botany. Nils accepted this offer. Blunt (2004), pp. 17–18. Stöver (1794), pp. 8–11.


University studies


Lund

Rothman showed Linnaeus that botany was a serious subject. He taught Linnaeus to classify plants according to Tournefort's system. Linnaeus was also taught about the sexual reproduction of plants, according to Sébastien Vaillant. In 1727, Linnaeus, age 21, enrolled in Lund University in
Skåne Scania, also known by its native name of Skåne (, ), is the southernmost of the historical provinces (''landskap'') of Sweden. Located in the south tip of the geographical region of Götaland, the province is roughly conterminous with Skåne C ...
. Blunt (2004), p. 18. Stöver (1794), p. 13. He was registered as ', the Latin form of his full name, which he also used later for his Latin publications. Professor Kilian Stobæus, natural scientist, physician and historian, offered Linnaeus tutoring and lodging, as well as the use of his library, which included many books about botany. He also gave the student free admission to his lectures. Blunt (2004), pp. 21–22. Stöver (1794), p. 15. In his spare time, Linnaeus explored the flora of Skåne, together with students sharing the same interests. Stöver (1794), pp. 14–15.


Uppsala

In August 1728, Linnaeus decided to attend Uppsala University on the advice of Rothman, who believed it would be a better choice if Linnaeus wanted to study both medicine and botany. Rothman based this recommendation on the two professors who taught at the medical faculty at Uppsala: Olof Rudbeck the Younger and Lars Roberg. Although Rudbeck and Roberg had undoubtedly been good professors, by then they were older and not so interested in teaching. Rudbeck no longer gave public lectures, and had others stand in for him. The botany, zoology, pharmacology and anatomy lectures were not in their best state. Blunt (2004), pp. 23–25. In Uppsala, Linnaeus met a new benefactor, Olof Celsius, who was a professor of theology and an amateur botanist. Blunt (2004), pp. 31–32. He received Linnaeus into his home and allowed him use of his library, which was one of the richest botanical libraries in Sweden. Stöver (1794), pp. 19–20. In 1729, Linnaeus wrote a thesis, ' on plant sexual reproduction. This attracted the attention of Rudbeck; in May 1730, he selected Linnaeus to give lectures at the University although the young man was only a second-year student. His lectures were popular, and Linnaeus often addressed an audience of 300 people. Blunt (2004), pp. 32–34. In June, Linnaeus moved from Celsius's house to Rudbeck's to become the tutor of the three youngest of his 24 children. His friendship with Celsius did not wane and they continued their botanical expeditions. Blunt (2004), pp. 34–37. Over that winter, Linnaeus began to doubt Tournefort's system of classification and decided to create one of his own. His plan was to divide the plants by the number of
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s and
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
s. He began writing several books, which would later result in, for example, ' and '. He also produced a book on the plants grown in the Uppsala Botanical Garden, '. Blunt (2001), pp. 36–37. Rudbeck's former assistant, Nils Rosén, returned to the University in March 1731 with a degree in medicine. Rosén started giving anatomy lectures and tried to take over Linnaeus's botany lectures, but Rudbeck prevented that. Until December, Rosén gave Linnaeus private tutoring in medicine. In December, Linnaeus had a "disagreement" with Rudbeck's wife and had to move out of his mentor's house; his relationship with Rudbeck did not appear to suffer. That Christmas, Linnaeus returned home to Stenbrohult to visit his parents for the first time in about three years. His mother had disapproved of his failing to become a priest, but she was pleased to learn he was teaching at the University. Anderson (1997), p. 40.


Expedition to Lapland

During a visit with his parents, Linnaeus told them about his plan to travel to
Lapland Lapland may refer to: Places *Lapland or Sápmi, an ethno-cultural region stretching over northern Fennoscandia (parts of Norway, Sweden, Finland, and Russia) **Lapland (Finland) (''Lappi''/''Lappland''), a Finnish region *** Lapland (former pr ...
; Rudbeck had made the journey in 1695, but the detailed results of his exploration were lost in a fire seven years afterwards. Linnaeus's hope was to find new plants, animals and possibly valuable minerals. He was also curious about the customs of the native
Sami people Acronyms * SAMI, ''Synchronized Accessible Media Interchange'', a closed-captioning format developed by Microsoft * Saudi Arabian Military Industries, a government-owned defence company * South African Malaria Initiative, a virtual expertise net ...
, reindeer-herding nomads who wandered Scandinavia's vast tundras. In April 1732, Linnaeus was awarded a grant from the Royal Society of Sciences in Uppsala for his journey. Anderson (1997), pp. 42–43. Blunt (2001), p. 38. Linnaeus began his expedition from Uppsala on 12 May 1732, just before he turned 25. He travelled on foot and horse, bringing with him his journal, botanical and ornithological manuscripts and sheets of paper for pressing plants. Near
Gävle Gävle () is a city in Sweden, the seat of Gävle Municipality and the capital of Gävleborg County. It had 77,586 inhabitants in 2020, which makes it the 13th most populated city in Sweden. It is the oldest city in the historical Norrland (Swede ...
he found great quantities of ''Campanula serpyllifolia'', later known as '' Linnaea borealis'', the twinflower that would become his favourite. Blunt (2001), pp. 42–43. He sometimes dismounted on the way to examine a flower or rock Anderson (1997), pp. 43–44. and was particularly interested in
moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
es and
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.reindeer, a common and economically important animal in Lapland. Anderson (1997), p. 46. Linnaeus travelled clockwise around the coast of the
Gulf of Bothnia The Gulf of Bothnia (; fi, Pohjanlahti; sv, Bottniska viken) is divided into the Bothnian Bay and Bothnian Sea, and it is the northernmost arm of the Baltic Sea, between Finland's west coast ( East Bothnia) and the Sweden's east coast (West ...
, making major inland incursions from Umeå,
Luleå Luleå ( , , locally ; smj, Luleju; fi, Luulaja) is a city on the coast of northern Sweden, and the capital of Norrbotten County, the northernmost county in Sweden. Luleå has 48,728 inhabitants in its urban core (2018) and is the seat of Lu ...
and Tornio. He returned from his six-month-long, over expedition in October, having gathered and observed many plants, birds and rocks. Blunt (2001), pp. 63–65. Blunt (2004), pp. 39–42. Broberg (2006), p. 29. Although Lapland was a region with limited biodiversity, Linnaeus described about 100 previously unidentified plants. These became the basis of his book '. Quammen (2007), p. 2. Stöver (1794), pp. 38–39. However, on the expedition to Lapland, Linnaeus used Latin names to describe organisms because he had not yet developed the binomial system. In ' Linnaeus's ideas about nomenclature and
classification Classification is a process related to categorization, the process in which ideas and objects are recognized, differentiated and understood. Classification is the grouping of related facts into classes. It may also refer to: Business, organizat ...
were first used in a practical way, making this the first proto-modern Flora. The account covered 534 species, used the Linnaean classification system and included, for the described species, geographical distribution and taxonomic notes. It was Augustin Pyramus de Candolle who attributed Linnaeus with ' as the first example in the botanical genre of Flora writing. Botanical historian E. L. Greene described ' as "the most classic and delightful" of Linnaeus's works. Frodin (2001), p. 27. It was also during this expedition that Linnaeus had a flash of insight regarding the classification of mammals. Upon observing the lower jawbone of a horse at the side of a road he was travelling, Linnaeus remarked: "If I only knew how many teeth and of what kind every animal had, how many teats and where they were placed, I should perhaps be able to work out a perfectly natural system for the arrangement of all quadrupeds." Blunt (2001), p. 54. In 1734, Linnaeus led a small group of students to Dalarna. Funded by the Governor of Dalarna, the expedition was to catalogue known natural resources and discover new ones, but also to gather intelligence on Norwegian mining activities at Røros.


Years in the Dutch Republic (1735–38)


Doctorate

His relations with Nils Rosén having worsened, Linnaeus accepted an invitation from Claes Sohlberg, son of a mining inspector, to spend the Christmas holiday in Falun, where Linnaeus was permitted to visit the mines. Blunt (2001), p. 74. In April 1735, at the suggestion of Sohlberg's father, Linnaeus and Sohlberg set out for the Dutch Republic, where Linnaeus intended to study medicine at the University of Harderwijk Stöver (1794), p. 71. while tutoring Sohlberg in exchange for an annual salary. At the time, it was common for Swedes to pursue doctoral degrees in the Netherlands, then a highly revered place to study natural history. Blunt (2001), pp. 78–79. On the way, the pair stopped in Hamburg, where they met the mayor, who proudly showed them a supposed wonder of nature in his possession: the taxidermied remains of a seven-headed
hydra Hydra generally refers to: * Lernaean Hydra, a many-headed serpent in Greek mythology * ''Hydra'' (genus), a genus of simple freshwater animals belonging to the phylum Cnidaria Hydra or The Hydra may also refer to: Astronomy * Hydra (constel ...
. Linnaeus quickly discovered the specimen was a
fake Fake may refer to: * Deception, an act or a statement intended to deceive ** Charlatan, a person who practices deception to obtain money or other advantages ** Counterfeit, a reproduction of an item, intended to deceive ** Cover-up, an attempt to ...
, cobbled together from the jaws and paws of weasels and the skins of snakes. The provenance of the hydra suggested to Linnaeus that it had been manufactured by monks to represent the
Beast of Revelation The Beast ( grc-x-koine, Θηρίον, ) may refer to one of two beasts described in the Book of Revelation. In Revelation 13:1-10, the first beast (interpreted as the Antichrist) rises "out of the sea" and is given authority and power by the ...
. Even at the risk of incurring the mayor's wrath, Linnaeus made his observations public, dashing the mayor's dreams of selling the hydra for an enormous sum. Linnaeus and Sohlberg were forced to flee from Hamburg. Anderson (1997), pp. 60–61. Blunt (2004), p. 90. Linnaeus began working towards his degree as soon as he reached Harderwijk, a university known for awarding degrees in as little as a week. He submitted a dissertation, written back in Sweden, entitled ''Dissertatio medica inauguralis in qua exhibetur hypothesis nova de febrium intermittentium causa'',That is, ''Inaugural thesis in medicine, in which a new hypothesis on the cause of intermittent fevers is presented'' in which he laid out his hypothesis that malaria arose only in areas with clay-rich soils. Although he failed to identify the true source of disease transmission, (i.e., the '' Anopheles'' mosquito), he did correctly predict that '' Artemisia annua'' (
wormwood Wormwood may refer to: Biology * Several plants of the genus ''Artemisia'': ** ''Artemisia abrotanum'', southern wormwood ** '' Artemisia absinthium'', common wormwood, grande wormwood or absinthe wormwood ** ''Artemisia annua'', sweet wormwood o ...
) would become a source of antimalarial medications. Within two weeks he had completed his oral and practical examinations and was awarded a doctoral degree. Blunt (2001)
p. 94
That summer Linnaeus reunited with Peter Artedi, a friend from Uppsala with whom he had once made a pact that should either of the two predecease the other, the survivor would finish the decedent's work. Ten weeks later, Artedi drowned in the
canals of Amsterdam Amsterdam, capital of the Netherlands, has more than of ''grachten'' (canals), about 90 islands and 1,500 bridges. The three main canals (Herengracht, Prinsengracht and Keizersgracht), dug in the 17th century during the Dutch Golden Age, form c ...
, leaving behind an unfinished manuscript on the classification of fish. Anderson (1997), p. 66. Blunt (2004), pp. 98–100.


Publishing of '

One of the first scientists Linnaeus met in the Netherlands was Johan Frederik Gronovius, to whom Linnaeus showed one of the several manuscripts he had brought with him from Sweden. The manuscript described a new system for classifying plants. When Gronovius saw it, he was very impressed, and offered to help pay for the printing. With an additional monetary contribution by the Scottish doctor Isaac Lawson, the manuscript was published as ' (1735). Blunt (2001), p. 98. Anderson (1997), pp. 62–63. Linnaeus became acquainted with one of the most respected physicians and botanists in the Netherlands, Herman Boerhaave, who tried to convince Linnaeus to make a career there. Boerhaave offered him a journey to South Africa and America, but Linnaeus declined, stating he would not stand the heat. Instead, Boerhaave convinced Linnaeus that he should visit the botanist Johannes Burman. After his visit, Burman, impressed with his guest's knowledge, decided Linnaeus should stay with him during the winter. During his stay, Linnaeus helped Burman with his '. Burman also helped Linnaeus with the books on which he was working: ' and '. Blunt (2004), pp. 100–102.


George Clifford, Philip Miller, and Johann Jacob Dillenius

In August 1735, during Linnaeus's stay with Burman, he met George Clifford III, a director of the Dutch East India Company and the owner of a rich botanical garden at the estate of Hartekamp in Heemstede. Clifford was very impressed with Linnaeus's ability to classify plants, and invited him to become his physician and superintendent of his garden. Linnaeus had already agreed to stay with Burman over the winter, and could thus not accept immediately. However, Clifford offered to compensate Burman by offering him a copy of Sir Hans Sloane's ''Natural History of Jamaica'', a rare book, if he let Linnaeus stay with him, and Burman accepted. Anderson (1997), p. 64. Stöver (1794), pp. 81–82. On 24 September 1735, Linnaeus moved to Hartekamp to become personal physician to Clifford, and curator of Clifford's herbarium. He was paid 1,000
florin The Florentine florin was a gold coin struck from 1252 to 1533 with no significant change in its design or metal content standard during that time. It had 54 grains (3.499 grams, 0.113 troy ounce) of nominally pure or 'fine' gold with a purcha ...
s a year, with free board and lodging. Though the agreement was only for a winter of that year, Linnaeus practically stayed there until 1738. It was here that he wrote a book ''Hortus Cliffortianus'', in the preface of which he described his experience as "the happiest time of my life". (A portion of Hartekamp was declared as public garden in April 1956 by the Heemstede local authority, and was named "Linnaeushof". It eventually became, as it is claimed, the biggest playground in Europe.) In July 1736, Linnaeus travelled to England, at Clifford's expense. Blunt (2001), pp. 106–107. He went to London to visit Sir Hans Sloane, a collector of natural history, and to see his cabinet, Stöver (1794), p. 89. as well as to visit the Chelsea Physic Garden and its keeper, Philip Miller. He taught Miller about his new system of subdividing plants, as described in '. Miller was in fact reluctant to use the new binomial nomenclature, preferring the classifications of Joseph Pitton de Tournefort and
John Ray John Ray FRS (29 November 1627 – 17 January 1705) was a Christian English naturalist widely regarded as one of the earliest of the English parson-naturalists. Until 1670, he wrote his name as John Wray. From then on, he used 'Ray', after ...
at first. Linnaeus, nevertheless, applauded Miller's ''Gardeners Dictionary'', The conservative Scot actually retained in his dictionary a number of pre-Linnaean binomial signifiers discarded by Linnaeus but which have been retained by modern botanists. He only fully changed to the Linnaean system in the edition of ''
The Gardeners Dictionary ''The Gardeners Dictionary'' was a widely cited reference series, written by Philip Miller (1691–1771), which tended to focus on plants cultivated in England. Eight editions of the series were published in his lifetime. After his death, it was ...
'' of 1768. Miller ultimately was impressed, and from then on started to arrange the garden according to Linnaeus's system. Stöver (1794), pp. 89–90. Linnaeus also travelled to Oxford University to visit the botanist Johann Jacob Dillenius. He failed to make Dillenius publicly fully accept his new classification system, though the two men remained in correspondence for many years afterwards. Linnaeus dedicated his ''Critica Botanica'' to him, as "''opus botanicum quo absolutius mundus non-vidit''". Linnaeus would later name a genus of tropical tree Dillenia in his honour. He then returned to Hartekamp, bringing with him many specimens of rare plants. Stöver (1794), pp. 90–93. The next year, 1737, he published ', in which he described 935
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
of plants, and shortly thereafter he supplemented it with ', with another sixty (''sexaginta'') genera. Stöver (1794), p. 95. His work at Hartekamp led to another book, ', a catalogue of the botanical holdings in the herbarium and botanical garden of Hartekamp. He wrote it in nine months (completed in July 1737), but it was not published until 1738. It contains the first use of the name ''
Nepenthes ''Nepenthes'' () is a genus of carnivorous plants, also known as tropical pitcher plants, or monkey cups, in the monotypic family Nepenthaceae. The genus includes about 170 species, and numerous natural and many cultivated hybrids. They are mos ...
'', which Linnaeus used to describe a genus of pitcher plants."If this is not Helen's '' Nepenthes'', it certainly will be for all botanists. What botanist would not be filled with admiration if, after a long journey, he should find this wonderful plant. In his astonishment past ills would be forgotten when beholding this admirable work of the Creator!" (translated from Latin by
Harry Veitch Sir Harry James Veitch (24 June 1840 – 6 July 1924) was an eminent English horticulturist in the nineteenth century, who was the head of the family nursery business, James Veitch & Sons, based in Chelsea, London. He was instrumental in esta ...
)
Linnaeus stayed with Clifford at Hartekamp until 18 October 1737 (new style), when he left the house to return to Sweden. Illness and the kindness of Dutch friends obliged him to stay some months longer in Holland. In May 1738, he set out for Sweden again. On the way home, he stayed in Paris for about a month, visiting botanists such as Antoine de Jussieu. After his return, Linnaeus never left Sweden again. Blunt (2001), p. 123. Koerner (1999), p. 56.


Return to Sweden

When Linnaeus returned to Sweden on 28 June 1738, he went to Falun, where he entered into an engagement to Sara Elisabeth Moræa. Three months later, he moved to Stockholm to find employment as a physician, and thus to make it possible to support a family. Stöver (1794), p. 141. Once again, Linnaeus found a patron; he became acquainted with Count Carl Gustav Tessin, who helped him get work as a physician at the Admiralty. Stöver (1794), pp. 146–147. Koerner (1999), p. 16. During this time in Stockholm, Linnaeus helped found the Royal Swedish Academy of Science; he became the first Praeses of the academy by drawing of lots. Koerner (1999), pp. 103–105. Because his finances had improved and were now sufficient to support a family, he received permission to marry his fiancée, Sara Elisabeth Moræa. Their wedding was held 26 June 1739. Seventeen months later, Sara gave birth to their first son, Carl. Two years later, a daughter, Elisabeth Christina, was born, and the subsequent year Sara gave birth to Sara Magdalena, who died when 15 days old. Sara and Linnaeus would later have four other children: Lovisa, , Johannes and Sophia. Stöver (1794), p. 382. In May 1741, Linnaeus was appointed Professor of Medicine at Uppsala University, first with responsibility for medicine-related matters. Soon, he changed place with the other Professor of Medicine, Nils Rosén, and thus was responsible for the Botanical Garden (which he would thoroughly reconstruct and expand), botany and natural history, instead. In October that same year, his wife and nine-month-old son followed him to live in Uppsala. Gribbin & Gribbin (2008), pp. 49–50.


Öland and Gotland

Ten days after he was appointed Professor, he undertook an expedition to the island provinces of Öland and
Gotland Gotland (, ; ''Gutland'' in Gutnish), also historically spelled Gottland or Gothland (), is Sweden's largest island. It is also a province, county, municipality, and diocese. The province includes the islands of Fårö and Gotska Sandön to the ...
with six students from the university to look for plants useful in medicine. First, they travelled to Öland and stayed there until 21 June, when they sailed to Visby in Gotland. Linnaeus and the students stayed on Gotland for about a month, and then returned to Uppsala. During this expedition, they found 100 previously unrecorded plants. The observations from the expedition were later published in ', written in Swedish. Like ', it contained both zoological and botanical observations, as well as observations concerning the culture in Öland and Gotland. Koerner (1999), p. 115. Blunt (2004), pp. 137–142. During the summer of 1745, Linnaeus published two more books: ' and '. ' was a strictly botanical book, while ' was zoological. Stöver (1794), pp. 117–118. Anders Celsius had created the temperature scale named after him in 1742. Celsius's scale was inverted compared to today, the boiling point at 0 °C and freezing point at 100 °C. In 1745, Linnaeus inverted the scale to its present standard. Koerner (1999), p. 204.


Västergötland

In the summer of 1746, Linnaeus was once again commissioned by the Government to carry out an expedition, this time to the Swedish province of Västergötland. He set out from Uppsala on 12 June and returned on 11 August. On the expedition his primary companion was Erik Gustaf Lidbeck, a student who had accompanied him on his previous journey. Linnaeus described his findings from the expedition in the book ', published the next year. Blunt (2004), p. 159. After he returned from the journey, the Government decided Linnaeus should take on another expedition to the southernmost province Scania. This journey was postponed, as Linnaeus felt too busy. In 1747, Linnaeus was given the title archiater, or chief physician, by the Swedish king
Adolf Frederick Adolf Frederick, or Adolph Frederick ( sv, Adolf Fredrik, german: Adolf Friedrich; 14 May 171012 February 1771) was King of Sweden from 1751 until his death. He was the son of Christian August of Holstein-Gottorp, Prince of Eutin, and Albertina ...
—a mark of great respect. Blunt (2004), p. 165. The same year he was elected member of the Academy of Sciences in Berlin. Stöver (1794), p. 167.


Scania

In the spring of 1749, Linnaeus could finally journey to Scania, again commissioned by the Government. With him he brought his student, Olof Söderberg. On the way to Scania, he made his last visit to his brothers and sisters in Stenbrohult since his father had died the previous year. The expedition was similar to the previous journeys in most aspects, but this time he was also ordered to find the best place to grow walnut and
Swedish whitebeam ''Scandosorbus intermedia'', the Swedish whitebeam, is a species of whitebeam found in southern Sweden, with scattered occurrences in easternmost Denmark (Bornholm), the far southwest of Finland, Estonia, Latvia and northern Poland.Rushforth, K. ...
trees; these trees were used by the military to make rifles. While there, they also visited the Ramlösa mineral spa, where he remarked on the quality of its ferruginous water. The journey was successful, and Linnaeus's observations were published the next year in '. Blunt (2004), pp. 198–205. Koerner (1999), p. 116.


Rector of Uppsala University

In 1750, Linnaeus became rector of Uppsala University, starting a period where natural sciences were esteemed. Perhaps the most important contribution he made during his time at Uppsala was to teach; many of his students travelled to various places in the world to collect botanical samples. Linnaeus called the best of these students his "apostles". Gribbin & Gribbin (2008), pp. 56–57. His lectures were normally very popular and were often held in the Botanical Garden. He tried to teach the students to think for themselves and not trust anybody, not even him. Even more popular than the lectures were the botanical excursions made every Saturday during summer, where Linnaeus and his students explored the flora and fauna in the vicinity of Uppsala. Blunt (2004), pp. 173–174.


''Philosophia Botanica''

Linnaeus published '' Philosophia Botanica'' in 1751. The book contained a complete survey of the taxonomy system he had been using in his earlier works. It also contained information of how to keep a journal on travels and how to maintain a botanical garden. Blunt (2004), p. 221.


''Nutrix Noverca''

During Linnaeus's time it was normal for upper class women to have wet nurses for their babies. Linnaeus joined an ongoing campaign to end this practice in Sweden and promote breast-feeding by mothers. In 1752 Linnaeus published a thesis along with Frederick Lindberg, a physician student, based on their experiences. In the tradition of the period, this dissertation was essentially an idea of the presiding reviewer (''prases'') expounded upon by the student. Linnaeus's dissertation was translated into French by J. E. Gilibert in 1770 as ''La Nourrice marâtre, ou Dissertation sur les suites funestes du nourrisage mercénaire''. Linnaeus suggested that children might absorb the personality of their wet nurse through the milk. He admired the child care practices of the Lapps and pointed out how healthy their babies were compared to those of Europeans who employed wet nurses. He compared the behaviour of wild animals and pointed out how none of them denied their newborns their breastmilk. It is thought that his activism played a role in his choice of the term ''Mammalia'' for the class of organisms.


''Species Plantarum''

Linnaeus published ''
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the ...
'', the work which is now internationally accepted as the starting point of modern
botanical nomenclature Botanical nomenclature is the formal, scientific naming of plants. It is related to, but distinct from Alpha taxonomy, taxonomy. Plant taxonomy is concerned with grouping and classifying plants; botanical nomenclature then provides names for the ...
, in 1753. Stace (1991)
p. 24
The first volume was issued on 24 May, the second volume followed on 16 August of the same year. The book contained 1,200 pages and was published in two volumes; it described over 7,300 species. Gribbin & Gribbin (2008), p. 47. Stöver (1794), pp. 198–199. The same year the king dubbed him knight of the Order of the Polar Star, the first civilian in Sweden to become a knight in this order. He was then seldom seen not wearing the order's insignia. Blunt (2004), p. 166.


Ennoblement

Linnaeus felt Uppsala was too noisy and unhealthy, so he bought two farms in 1758: Hammarby and Sävja. The next year, he bought a neighbouring farm, Edeby. He spent the summers with his family at Hammarby; initially it only had a small one-storey house, but in 1762 a new, larger main building was added. Blunt (2004), p. 219. In Hammarby, Linnaeus made a garden where he could grow plants that could not be grown in the Botanical Garden in Uppsala. He began constructing a museum on a hill behind Hammarby in 1766, where he moved his library and collection of plants. A fire that destroyed about one third of Uppsala and had threatened his residence there necessitated the move. Blunt (2004), pp. 220–224. Since the initial release of ' in 1735, the book had been expanded and reprinted several times; the tenth edition was released in 1758. This edition established itself as the starting point for zoological nomenclature, the equivalent of '. Blunt (2004), p. 6. The Swedish King Adolf Frederick granted Linnaeus nobility in 1757, but he was not ennobled until 1761. With his ennoblement, he took the name Carl von Linné (Latinised as '), 'Linné' being a shortened and gallicised version of 'Linnæus', and the German
nobiliary particle A nobiliary particle is used in a surname or family name in many Western cultures to signal the nobility of a family. The particle used varies depending on the country, language and period of time. In some languages, it is the same as a regular p ...
' von' signifying his ennoblement. The noble family's coat of arms prominently features a twinflower, one of Linnaeus's favourite plants; it was given the scientific name ''Linnaea borealis'' in his honour by Gronovius. The shield in the coat of arms is divided into thirds: red, black and green for the three kingdoms of nature (animal, mineral and vegetable) in Linnaean classification; in the centre is an egg "to denote Nature, which is continued and perpetuated ''in ovo''." At the bottom is a phrase in Latin, borrowed from the Aeneid, which reads "''Famam extendere factis''": we extend our fame by our deeds. Gribbin & Gribbin (2008), p. 62. Blunt (2004), p. 199. Blunt (2004), pp. 229–230. Linnaeus inscribed this personal motto in books that were given to him by friends. After his ennoblement, Linnaeus continued teaching and writing. His reputation had spread over the world, and he corresponded with many different people. For example,
Catherine II of Russia , en, Catherine Alexeievna Romanova, link=yes , house = , father = Christian August, Prince of Anhalt-Zerbst , mother = Joanna Elisabeth of Holstein-Gottorp , birth_date = , birth_name = Princess Sophie of Anhal ...
sent him seeds from her country.Uppsala University, Linné Online
English language version
He also corresponded with Giovanni Antonio Scopoli, "the Linnaeus of the Austrian Empire", who was a doctor and a botanist in Idrija, Duchy of Carniola (nowadays Slovenia). Scopoli communicated all of his research, findings, and descriptions (for example of the olm and the dormouse, two little animals hitherto unknown to Linnaeus). Linnaeus greatly respected Scopoli and showed great interest in his work. He named a solanaceous genus, '' Scopolia'', the source of
scopolamine Scopolamine, also known as hyoscine, or Devil's Breath, is a natural or synthetically produced tropane alkaloid and anticholinergic drug that is formally used as a medication for treating motion sickness and postoperative nausea and vomiting ...
, after him, but because of the great distance between them, they never met.


Final years

Linnaeus was relieved of his duties in the Royal Swedish Academy of Science in 1763, but continued his work there as usual for more than ten years after. In 1769 he was elected to the American Philosophical Society for his work. He stepped down as rector at Uppsala University in December 1772, mostly due to his declining health. Blunt (2004), p. 245. Linnaeus's last years were troubled by illness. He had had a disease called the Uppsala fever in 1764, but survived due to the care of Rosén. He developed sciatica in 1773, and the next year, he had a stroke which partially paralysed him. Blunt (2004), p. 232. He had a second stroke in 1776, losing the use of his right side and leaving him bereft of his memory; while still able to admire his own writings, he could not recognise himself as their author. Stöver (1794), pp. 243–245. Broberg (2006), p. 42. In December 1777, he had another stroke which greatly weakened him, and eventually led to his death on 10 January 1778 in Hammarby. Gribbin & Gribbin (2008), p. 63. Despite his desire to be buried in Hammarby, he was buried in Uppsala Cathedral on 22 January. Quammen (2007), p. 4. Anderson (1997), pp. 104–106. His library and collections were left to his widow Sara and their children.
Joseph Banks Sir Joseph Banks, 1st Baronet, (19 June 1820) was an English naturalist, botanist, and patron of the natural sciences. Banks made his name on the 1766 natural-history expedition to Newfoundland and Labrador. He took part in Captain James ...
, an eminent botanist, wished to purchase the collection, but his son Carl refused the offer and instead moved the collection to Uppsala. In 1783 Carl died and Sara inherited the collection, having outlived both her husband and son. She tried to sell it to Banks, but he was no longer interested; instead an acquaintance of his agreed to buy the collection. The acquaintance was a 24-year-old medical student, James Edward Smith, who bought the whole collection: 14,000 plants, 3,198 insects, 1,564 shells, about 3,000 letters and 1,600 books. Smith founded the Linnean Society of London five years later. Blunt (2001), pp. 238–240. The von Linné name ended with his son Carl, who never married. His other son, Johannes, had died aged 3. There are over two hundred descendants of Linnaeus through two of his daughters.


Apostles

During Linnaeus's time as Professor and Rector of Uppsala University, he taught many devoted students, 17 of whom he called "apostles". They were the most promising, most committed students, and all of them made botanical expeditions to various places in the world, often with his help. The amount of this help varied; sometimes he used his influence as Rector to grant his apostles a scholarship or a place on an expedition. Blunt (2004), pp. 189–190. To most of the apostles he gave instructions of what to look for on their journeys. Abroad, the apostles collected and organised new plants, animals and minerals according to Linnaeus's system. Most of them also gave some of their collection to Linnaeus when their journey was finished. Broberg (2006), pp. 37–39. Thanks to these students, the Linnaean system of taxonomy spread through the world without Linnaeus ever having to travel outside Sweden after his return from Holland. Anderson (1997), pp. 92–93. The British botanist William T. Stearn notes, without Linnaeus's new system, it would not have been possible for the apostles to collect and organise so many new specimens. Blunt (2004), pp. 184–185. Many of the apostles died during their expeditions.


Early expeditions

Christopher Tärnström, the first apostle and a 43-year-old pastor with a wife and children, made his journey in 1746. He boarded a Swedish East India Company ship headed for China. Tärnström never reached his destination, dying of a tropical fever on
Côn Sơn Island Côn Sơn ( ), also known as Côn Lôn is the largest island of the Côn Đảo archipelago, off the coast of southern Vietnam.Kelley, p 116 Other names Its French variant Grande-Condore was well-known during the times of French Indochina. Mar ...
the same year. Tärnström's widow blamed Linnaeus for making her children fatherless, causing Linnaeus to prefer sending out younger, unmarried students after Tärnström. Blunt (2004), pp. 185–186. Six other apostles later died on their expeditions, including Pehr Forsskål and Pehr Löfling. Two years after Tärnström's expedition, Finnish-born Pehr Kalm set out as the second apostle to North America. There he spent two-and-a-half years studying the flora and fauna of Pennsylvania, New York, New Jersey and Canada. Linnaeus was overjoyed when Kalm returned, bringing back with him many pressed flowers and seeds. At least 90 of the 700 North American species described in ''Species Plantarum'' had been brought back by Kalm. Anderson (1997), pp. 93–94.


Cook expeditions and Japan

Daniel Solander was living in Linnaeus's house during his time as a student in Uppsala. Linnaeus was very fond of him, promising Solander his eldest daughter's hand in marriage. On Linnaeus's recommendation, Solander travelled to England in 1760, where he met the English botanist
Joseph Banks Sir Joseph Banks, 1st Baronet, (19 June 1820) was an English naturalist, botanist, and patron of the natural sciences. Banks made his name on the 1766 natural-history expedition to Newfoundland and Labrador. He took part in Captain James ...
. With Banks, Solander joined
James Cook James Cook (7 November 1728 Old Style date: 27 October – 14 February 1779) was a British explorer, navigator, cartographer, and captain in the British Royal Navy, famous for his three voyages between 1768 and 1779 in the Pacific Ocean an ...
on his expedition to Oceania on the ''
Endeavour Endeavour or endeavor may refer to: People Fictional characters * Endeavour Morse, central character of the ''Inspector Morse'' novels by Colin Dexter * Endeavor, the hero name for the character Enji Todoroki from the anime series ''My Hero A ...
'' in 1768–71. Anderson (1997), p. 96. Blunt (2004), pp. 191–192. Solander was not the only apostle to journey with James Cook; Anders Sparrman followed on the '' Resolution'' in 1772–75 bound for, among other places, Oceania and South America. Sparrman made many other expeditions, one of them to South Africa. Blunt (2004), pp. 192–193. Perhaps the most famous and successful apostle was
Carl Peter Thunberg Carl Peter Thunberg, also known as Karl Peter von Thunberg, Carl Pehr Thunberg, or Carl Per Thunberg (11 November 1743 – 8 August 1828), was a Swedish naturalist and an "apostle" of Carl Linnaeus. After studying under Linnaeus at Uppsala Un ...
, who embarked on a nine-year expedition in 1770. He stayed in South Africa for three years, then travelled to
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
. All foreigners in Japan were forced to stay on the island of Dejima outside Nagasaki, so it was thus hard for Thunberg to study the flora. He did, however, manage to persuade some of the translators to bring him different plants, and he also found plants in the gardens of Dejima. He returned to Sweden in 1779, one year after Linnaeus's death. Blunt (2004), pp. 193–194.


Major publications


''Systema Naturae''

The first edition of ' was printed in the Netherlands in 1735. It was a twelve-page work. Linnaeus (1735) By the time it reached its
10th edition 1 (one, unit, unity) is a number representing a single or the only entity. 1 is also a numerical digit and represents a single unit of counting or measurement. For example, a line segment of ''unit length'' is a line segment of length 1. I ...
in 1758, it classified 4,400 species of animals and 7,700 species of plants. People from all over the world sent their specimens to Linnaeus to be included. By the time he started work on the 12th edition, Linnaeus needed a new invention—the
index card An index card (or record card in British English and system cards in Australian English) consists of card stock (heavy paper) cut to a standard size, used for recording and storing small amounts of discrete data. A collection of such cards e ...
—to track classifications. In ''Systema Naturae'', the unwieldy names mostly used at the time, such as "'", were supplemented with concise and now familiar "binomials", composed of the generic name, followed by a specific epithet—in the case given, '' Physalis angulata''. These binomials could serve as a label to refer to the species. Higher taxa were constructed and arranged in a simple and orderly manner. Although the system, now known as binomial nomenclature, was partially developed by the Bauhin brothers (see Gaspard Bauhin and Johann Bauhin) almost 200 years earlier, Linnaeus was the first to use it consistently throughout the work, including in monospecific genera, and may be said to have popularised it within the scientific community. After the decline in Linnaeus's health in the early 1770s, publication of editions of ''Systema Naturae'' went in two different directions. Another Swedish scientist, Johan Andreas Murray issued the ''Regnum Vegetabile'' section separately in 1774 as the ''Systema Vegetabilium'', rather confusingly labelled the 13th edition. Meanwhile, a 13th edition of the entire ''Systema'' appeared in parts between 1788 and 1793 under the editorship of
Johann Friedrich Gmelin , fields = , workplaces = University of GöttingenUniversity of Tübingen , alma_mater = University of Tübingen , doctoral_advisor = Philipp Friedrich GmelinFerdinand Christoph Oetinger , academic_advisors = , doctora ...
. It was through the ''Systema Vegetabilium'' that Linnaeus's work became widely known in England, following its translation from the Latin by the
Lichfield Botanical Society Erasmus Robert Darwin (12 December 173118 April 1802) was an English physician. One of the key thinkers of the Midlands Enlightenment, he was also a natural philosopher, physiologist, slave-trade abolitionist, inventor, and poet. His poems ...
as ''A System of Vegetables'' (1783–1785).


''Orbis eruditi judicium de Caroli Linnaei MD scriptis''

('Opinion of the learned world on the writings of Carl Linnaeus, Doctor') Published in 1740, this small octavo-sized pamphlet was presented to the State Library of New South Wales by the Linnean Society of NSW in 2018. This is considered among the rarest of all the writings of Linnaeus, and crucial to his career, securing him his appointment to a professorship of medicine at Uppsala University. From this position he laid the groundwork for his radical new theory of classifying and naming organisms for which he was considered the founder of modern taxonomy.


'

' (or, more fully, ') was first published in 1753, as a two-volume work. Its prime importance is perhaps that it is the primary starting point of
plant nomenclature Botanical nomenclature is the formal, scientific naming of plants. It is related to, but distinct from taxonomy. Plant taxonomy is concerned with grouping and classifying plants; botanical nomenclature then provides names for the results of this ...
as it exists today.


'

' was first published in 1737, delineating plant genera. Around 10 editions were published, not all of them by Linnaeus himself; the most important is the 1754 fifth edition. Stace (1991)
p. 22
In it Linnaeus divided the plant Kingdom into 24 classes. One, Cryptogamia, included all the plants with concealed reproductive parts (algae, fungi, mosses and liverworts and ferns). Van den Hoek ''et al.'' (2005).


'

' (1751) was a summary of Linnaeus's thinking on plant classification and nomenclature, and an elaboration of the work he had previously published in ' (1736) and ' (1737). Other publications forming part of his plan to reform the foundations of botany include his ' and ': all were printed in Holland (as were ' (1737) and ' (1735)), the ''Philosophia'' being simultaneously released in Stockholm.


Collections

At the end of his lifetime the Linnean collection in Uppsala was considered one of the finest collections of natural history objects in Sweden. Next to his own collection he had also built up a museum for the university of Uppsala, which was supplied by material donated by Carl Gyllenborg (in 1744–1745), crown-prince
Adolf Fredrik Adolf Frederick, or Adolph Frederick ( sv, Adolf Fredrik, german: Adolf Friedrich; 14 May 171012 February 1771) was King of Sweden from 1751 until his death. He was the son of Christian August of Holstein-Gottorp, Prince of Eutin, and Albertin ...
(in 1745), Erik Petreus (in 1746), Claes Grill (in 1746), Magnus Lagerström (in 1748 and 1750) and
Jonas Alströmer Jonas Alströmer (7 January 1685 – 2 June 1761) was a pioneer of agriculture and industry in Sweden. Born Jonas Toresson (later changed to Alström) in the town of Alingsås in Västergötland, in 1707 he became a clerk for Stockholm merchant ...
(in 1749). The relation between the museum and the private collection was not formalised and the steady flow of material from Linnean pupils were incorporated to the private collection rather than to the museum.Wallin, L. 2001.
Catalogue of type specimens. 4. Linnaean specimens. – pp. 1–128. Uppsala. (Uppsala University, Museum of Evolution, Zoology Section).
Linnaeus felt his work was reflecting the harmony of nature and he said in 1754 "the earth is then nothing else but a museum of the all-wise creator's masterpieces, divided into three chambers". He had turned his own estate into a microcosm of that 'world museum'. In April 1766 parts of the town were destroyed by a fire and the Linnean private collection was subsequently moved to a barn outside the town, and shortly afterwards to a single-room stone building close to his country house at Hammarby near Uppsala. This resulted in a physical separation between the two collections; the museum collection remained in the botanical garden of the university. Some material which needed special care (alcohol specimens) or ample storage space was moved from the private collection to the museum. In Hammarby the Linnean private collections suffered seriously from damp and the depredations by mice and insects. Carl von Linné's son (Carl Linnaeus) inherited the collections in 1778 and retained them until his own death in 1783. Shortly after Carl von Linné's death his son confirmed that mice had caused "horrible damage" to the plants and that also moths and mould had caused considerable damage.Dance, S.P. 1967. Report on the Linnaean shell collection. – Proceedings of the Linnean Society of London 178 (1): 1–24, Pl. 1–10. He tried to rescue them from the neglect they had suffered during his father's later years, and also added further specimens. This last activity however reduced rather than augmented the scientific value of the original material. In 1784 the young medical student James Edward Smith purchased the entire specimen collection, library, manuscripts, and correspondence of Carl Linnaeus from his widow and daughter and transferred the collections to London. Fries (2011), pp. 342–357. Not all material in Linné's private collection was transported to England. Thirty-three fish specimens preserved in alcohol were not sent and were later lost. In London Smith tended to neglect the zoological parts of the collection; he added some specimens and also gave some specimens away. Over the following centuries the Linnean collection in London suffered enormously at the hands of scientists who studied the collection, and in the process disturbed the original arrangement and labels, added specimens that did not belong to the original series and withdrew precious original type material. Much material which had been intensively studied by Linné in his scientific career belonged to the collection of Queen Lovisa Ulrika (1720–1782) (in the Linnean publications referred to as "Museum Ludovicae Ulricae" or "M. L. U."). This collection was donated by her grandson King Gustav IV Adolf (1778–1837) to the museum in Uppsala in 1804. Another important collection in this respect was that of her husband King Adolf Fredrik (1710–1771) (in the Linnean sources known as "Museum Adolphi Friderici" or "Mus. Ad. Fr."), the wet parts (alcohol collection) of which were later donated to the
Royal Swedish Academy of Sciences The Royal Swedish Academy of Sciences ( sv, Kungliga Vetenskapsakademien) is one of the Swedish Royal Academies, royal academies of Sweden. Founded on 2 June 1739, it is an independent, non-governmental scientific organization that takes special ...
, and is today housed in the
Swedish Museum of Natural History The Swedish Museum of Natural History ( sv, Naturhistoriska riksmuseet, literally, the National Museum of Natural History), in Stockholm, is one of two major museums of natural history in Sweden, the other one being located in Gothenburg. The ...
at
Stockholm Stockholm () is the Capital city, capital and List of urban areas in Sweden by population, largest city of Sweden as well as the List of urban areas in the Nordic countries, largest urban area in Scandinavia. Approximately 980,000 people liv ...
. The dry material was transferred to Uppsala.


System of taxonomy

The establishment of universally accepted conventions for the naming of organisms was Linnaeus's main contribution to taxonomy—his work marks the starting point of consistent use of binomial nomenclature. Reveal & Pringle (1993), pp. 160–161. During the 18th century expansion of natural history knowledge, Linnaeus also developed what became known as the ''Linnaean taxonomy''; the system of scientific classification now widely used in the biological sciences. A previous zoologist
Rumphius Georg Eberhard Rumphius (originally: Rumpf; baptized c. 1 November 1627 – 15 June 1702) was a Germans, German-born botanist employed by the Dutch East India Company in what is now eastern Indonesia, and is best known for his work ''Herbarium Am ...
(1627–1702) had more or less approximated the Linnaean system and his material contributed to the later development of the binomial scientific classification by Linnaeus. The Linnaean system classified nature within a nested hierarchy, starting with three kingdoms. Kingdoms were divided into classes and they, in turn, into orders, and thence into genera (''singular:'' genus), which were divided into species (''singular:'' species). Below the rank of species he sometimes recognised taxa of a lower (unnamed) rank; these have since acquired standardised names such as ''variety'' in botany and ''subspecies'' in zoology. Modern taxonomy includes a rank of family between order and genus and a rank of
phylum In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature f ...
between kingdom and class that were not present in Linnaeus's original system. Davis & Heywood (1973), p. 17. Linnaeus's groupings were based upon shared physical characteristics, and not based upon differences. Of his higher groupings, only those for animals are still in use, and the groupings themselves have been significantly changed since their conception, as have the principles behind them. Nevertheless, Linnaeus is credited with establishing the idea of a hierarchical structure of classification which is based upon observable characteristics and intended to reflect natural relationships. While the underlying details concerning what are considered to be scientifically valid "observable characteristics" have changed with expanding knowledge (for example,
DNA sequencing DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...
, unavailable in Linnaeus's time, has proven to be a tool of considerable utility for classifying living organisms and establishing their evolutionary relationships), the fundamental principle remains sound.


Human taxonomy

Linnaeus's system of taxonomy was especially noted as the first to include humans ('' Homo'') taxonomically grouped with apes (''
Simia In his ''Systema Naturae'' of 1758, Carl Linnaeus divided the Order Primates within '' Mammalia'' into four genera: '' Homo'', ''Simia'', '' Lemur'', and '' Vespertilio''. His ''Vespertilio'' included all bats, and has since been moved from Pri ...
''), under the header of '' Anthropomorpha''. German biologist
Ernst Haeckel Ernst Heinrich Philipp August Haeckel (; 16 February 1834 – 9 August 1919) was a German zoologist, naturalist, eugenicist, philosopher, physician, professor, marine biologist and artist. He discovered, described and named thousands of new sp ...
speaking in 1907 noted this as the "most important sign of Linnaeus's genius". Linnaeus classified humans among the primates beginning with the first edition of '. During his time at Hartekamp, he had the opportunity to examine several
monkey Monkey is a common name that may refer to most mammals of the infraorder Simiiformes, also known as the simians. Traditionally, all animals in the group now known as simians are counted as monkeys except the apes, which constitutes an incomple ...
s and noted similarities between them and man. Gribbin & Gribbin (2008), pp. 173–174. He pointed out both species basically have the same anatomy; except for speech, he found no other differences. Frängsmyr ''et al.'' (1983), p. 170. Frängsmyr ''et al.'' (1983), p. 167, quotes Linnaeus explaining the real difference would necessarily be absent from his classification system, as it was not a morphological characteristic: ''"I well know what a splendidly great difference there is etweena man and a ''bestia'' iterally, "beast"; that is, a non-human animalwhen I look at them from a point of view of morality. Man is the animal which the Creator has seen fit to honor with such a magnificent
mind The mind is the set of faculties responsible for all mental phenomena. Often the term is also identified with the phenomena themselves. These faculties include thought, imagination, memory, will, and sensation. They are responsible for various m ...
and has condescended to adopt as his favorite and for which he has prepared a nobler life"''. See als
books.google.com
in which Linnaeus cites the significant capacity to reason as the distinguishing characteristic of humans.
Thus he placed man and monkeys under the same category, ''Anthropomorpha'', meaning "manlike." Frängsmyr ''et al.'' (1983), p. 167. This classification received criticism from other biologists such as
Johan Gottschalk Wallerius Johan Gottschalk Wallerius (11 July 1709 – 16 November 1785) was a Swedish chemist and mineralogist. Biography Wallerius was born at Stora Mellösa in Närke (now Örebro County), Sweden. He was a son of provost Erik Nilsson Wallerius and h ...
,
Jacob Theodor Klein Jacob Theodor Klein (nickname ''Plinius Gedanensium''; 15 August 1685 – 27 February 1759) was a German jurist, historian, botanist, zoologist, mathematician and diplomat in service of Polish King August II the Strong. Life Klein was born on 15 ...
and Johann Georg Gmelin on the ground that it is illogical to describe man as human-like. In a letter to Gmelin from 1747, Linnaeus replied: The theological concerns were twofold: first, putting man at the same level as monkeys or apes would lower the spiritually higher position that man was assumed to have in the great chain of being, and second, because the Bible says man was created in the image of God ( theomorphism), if monkeys/apes and humans were not distinctly and separately designed, that would mean monkeys and apes were created in the image of God as well. This was something many could not accept. Frängsmyr ''et al.'' (1983), pp. 171–172. The conflict between world views that was caused by asserting man was a type of animal would simmer for a century until the much greater, and still ongoing, creation–evolution controversy began in earnest with the publication of '' On the Origin of Species'' by Charles Darwin in 1859. After such criticism, Linnaeus felt he needed to explain himself more clearly. The 10th edition of ' introduced new terms, including ''Mammalia'' and ''Primates'', the latter of which would replace ''Anthropomorpha'' Frängsmyr ''et al.'' (1983), p. 175. as well as giving humans the full binomial ''Homo sapiens''. Blunt (2004), p. 8. The new classification received less criticism, but many natural historians still believed he had demoted humans from their former place of ruling over nature and not being a part of it. Linnaeus believed that man biologically belongs to the animal kingdom and had to be included in it. Frängsmyr ''et al.'' (1983), pp. 191–192. In his book ', he said, "One should not vent one's wrath on animals, Theology decree that man has a soul and that the animals are mere 'aoutomata mechanica,' but I believe they would be better advised that animals have a soul and that the difference is of nobility." Frängsmyr ''et al.'' (1983), p. 166. Linnaeus added a second species to the genus ''Homo'' in ' based on a figure and description by Jacobus Bontius from a 1658 publication: ''Homo troglodytes'' ("caveman") Bontius (1658)
p. 84
and published a third in 1771: ''Homo lar''. Swedish historian Gunnar Broberg states that the new human species Linnaeus described were actually simians or native people clad in skins to frighten colonial settlers, whose appearance had been exaggerated in accounts to Linnaeus. Frängsmyr ''et al.'' (1983), p. 187. For ''Homo troglodytes'' Linnaeus asked the Swedish East India Company to search for one, but they did not find any signs of its existence. Frängsmyr ''et al.'' (1983), p. 186. ''Homo lar'' has since been reclassified as ''Hylobates lar'', the lar gibbon. In the first edition of ', Linnaeus subdivided the human species into four varieties: "Europæus albesc ns (whitish European), "Americanus rubesc ns (reddish American), "Asiaticus fuscus" (tawny Asian) and "Africanus nigr
culus Latin obscenity is the profane, indecent, or impolite vocabulary of Latin, and its uses. Words deemed obscene were described as (obscene, lewd, unfit for public use), or (improper, in poor taste, undignified). Documented obscenities occurred ra ...
(blackish African). In the tenth edition of Systema Naturae he further detailed phenotypical characteristics for each variety, based on the concept of the
four temperaments The four temperament theory is a proto-psychological theory which suggests that there are four fundamental personality types: sanguine, choleric, melancholic, and phlegmatic. Most formulations include the possibility of mixtures among the types w ...
from classical antiquity, and changed the description of Asians' skin tone to "luridus" (yellow). Additionally, Linnaeus created a wastebasket taxon "monstrosus" for "wild and monstrous humans, unknown groups, and more or less abnormal people". In 1959, W. T. Stearn designated Linnaeus to be the lectotype of ''H. sapiens''.


Influences and economic beliefs

Linnaeus's applied science was inspired not only by the instrumental utilitarianism general to the early Enlightenment, but also by his adherence to the older economic doctrine of Cameralism. Koerner (1999), p. 95–96. Additionally, Linnaeus was a state interventionist. He supported tariffs, levies, export bounties, quotas, embargoes, navigation acts, subsidised investment capital, ceilings on wages, cash grants, state-licensed producer monopolies, and cartels. Koerner (1999), p. 97.


Commemoration

Anniversaries of Linnaeus's birth, especially in centennial years, have been marked by major celebrations. Östholm (2007) Linnaeus has appeared on numerous Swedish postage stamps and banknotes. There are numerous statues of Linnaeus in countries around the world. The Linnean Society of London has awarded the
Linnean Medal The Linnean Medal of the Linnean Society of London was established in 1888, and is awarded annually to alternately a botanist or a zoologist or (as has been common since 1958) to one of each in the same year. The medal was of gold until 1976, and ...
for excellence in botany or zoology since 1888. Following approval by the Riksdag of Sweden, Växjö University and
Kalmar College Linnaeus University (LNU) ( sv, Linnéuniversitetet) is a state university in the Swedish historical province (''landskap'') Småland, with two campuses located in Växjö and Kalmar respectively. Linnaeus University was established in 2010 ...
merged on 1 January 2010 to become Linnaeus University. Other things named after Linnaeus include the twinflower genus '' Linnaea'', ''
Linnaeosicyos ''Linnaeosicyos'' is a monotypic genus of flowering plants belonging to the family Cucurbitaceae. The only species is ''Linnaeosicyos amara'' . It is native to the Dominican Republic. The genus name of ''Linnaeosicyos'' is in honour of Carl Linn ...
'' (a monotypic genus in the family Cucurbitaceae), the crater
Linné Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
on the Earth's moon, a street in Cambridge, Massachusetts, and the cobalt sulfide mineral Linnaeite.


Commentary

Andrew Dickson White wrote in '' A History of the Warfare of Science with Theology in Christendom'' (1896): The mathematical PageRank algorithm, applied to 24 multilingual Wikipedia editions in 2014, published in '' PLOS ONE'' in 2015, placed Carl Linnaeus at the top historical figure, above Jesus, Aristotle,
Napoleon Napoleon Bonaparte ; it, Napoleone Bonaparte, ; co, Napulione Buonaparte. (born Napoleone Buonaparte; 15 August 1769 – 5 May 1821), later known by his regnal name Napoleon I, was a French military commander and political leader who ...
, and Adolf Hitler (in that order). In the 21st century, Linnæus's taxonomy of human "races" has been problematised and discussed. Some critics claim that Linnæus was one of the forebears of the modern pseudoscientific notion of scientific racism, while others hold the view that while his classification was stereotyped, it did not imply that certain human "races" were superior to others.


Standard author abbreviation


Selected publications by Linnaeus

* * * * Linnaeus, Carl 1846 ''Fauna svecica. Sistens Animalia Sveciae Regni: Quadrupedia, Aves, Amphibia, Pisces, Insecta, Vermes, distributae per classes & ordines, genera & species''. C. Wishoff & G.J. Wishoff, Lugdnuni Batavorum. * * ''see also
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the ...
'' * * * * *


See also

* Linnaeus's flower clock * Johann Bartsch, colleague * ''
Centuria Insectorum The first page of ''Centuria Insectorum'', as included in ''Amoenitates Academicæ'' ' (Latin, "one hundred insects") is a 1763 taxonomic work by Carl Linnaeus, and defended as a thesis by Boas Johansson; which of the two men should for taxonomi ...
'' * History of botany *
History of phycology The history of phycology is the history of the scientific study of algae. Human interest in plants as food goes back into the origins of the species, and knowledge of algae can be traced back more than two thousand years. However, only in the last ...
* Scientific revolution


References


Notes


Citations


Sources

* * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * *


Further reading

* * * * * * * * *


External links

Biographies
Biography
at the Department of Systematic Botany, University of Uppsala
Biography
at The Linnean Society of London

from the University of California Museum of Paleontology * A four-minut
biographical video
from the London Natural History Museum on YouTube
Biography
from Taxonomic Literature, 2nd Edition. 1976–2009. Resources * *
The Linnean Society of London

The Linnaeus Apostles

The Linnean Collections

The Linnean Correspondence



The Linnaean Dissertations



The Linnaeus Tercentenary

Works by Carl von Linné
at the
Biodiversity Heritage Library The Biodiversity Heritage Library (BHL) is the world’s largest open access digital library for biodiversity literature and archives. BHL operates as worldwide consortiumof natural history, botanical, research, and national libraries working toge ...

Digital edition: "Critica Botanica"
by the
University and State Library Düsseldorf The University and State Library Düsseldorf (german: Universitäts- und Landesbibliothek Düsseldorf, abbreviated ULB Düsseldorf) is a central service institution of Heinrich Heine University. Along with Bonn and Münster, it is also one of th ...

Digital edition: "Classes plantarum seu systemata plantarum"
by the
University and State Library Düsseldorf The University and State Library Düsseldorf (german: Universitäts- und Landesbibliothek Düsseldorf, abbreviated ULB Düsseldorf) is a central service institution of Heinrich Heine University. Along with Bonn and Münster, it is also one of th ...

''Oratio de telluris habitabilis incremento''
(1744) – full digital facsimile from Linda Hall Library Other * Linnaeus was depicted by
Jay Hosler Jay Hosler is the author and illustrator of science-oriented comics. He is best known for his graphic novels '' Clan Apis'', '' The Sandwalk Adventures'', and '' Optical Allusions''. '' Clan Apis'', a Xeric Foundation Award winner, follows the li ...
in a parody of ''Peanuts'' titled
Good ol' Charlie Darwin
. * Th

of '' Nature'' featured a picture of Linnaeus on the cover with the heading "Linnaeus's Legacy" and devoted a substantial portion to items related to Linnaeus and Linnaean taxonomy. * A tattoo of Linnaeus'
definition of the order Primates
mentioned by Carl Zimmer * '' Ginkgo biloba'
tree at the University of Harderwijk
said to have been planted by Linnaeus in 1735 *''SL Magazine,'' Spring 2018 features an article by Nicholas Sparks, librarian, Collection Strategy and Development titled ''Origins of Taxonomy,'' describing a generous donation from the Linnean Society of NSW to supplement the State Library of New South Wales's collections on Carl Linnaeus of documents, photographs, prints and drawings as well as a fine portrait of Linnaeus painted about 1800. {{DEFAULTSORT:Linnaeus, Carl 1707 births 1778 deaths 18th-century Latin-language writers 18th-century male writers 18th-century Swedish physicians 18th-century Swedish zoologists 18th-century Swedish writers Age of Liberty people Swedish arachnologists Botanical nomenclature Botanists active in Europe Botanists with author abbreviations Bryologists Burials at Uppsala Cathedral Fellows of the Royal Society Historical definitions of race Knights of the Order of the Polar Star Members of the French Academy of Sciences Members of the Prussian Academy of Sciences Members of the Royal Swedish Academy of Sciences People from Älmhult Municipality Phycologists Pteridologists Swedish autobiographers Swedish biologists Swedish botanists Swedish entomologists Swedish expatriates in the Dutch Republic Swedish Lutherans Swedish mammalogists Swedish mycologists Linne, Carl von Swedish ornithologists Swedish taxonomists Terminologists Taxon authorities of Hypericum species University of Harderwijk alumni Uppsala University alumni Uppsala University faculty Members of the American Philosophical Society 18th-century lexicographers