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The kunga was a
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
equid Equidae (sometimes known as the horse family) is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. All extant species are in the genus '' Equus'', w ...
that was used as a draft animal in ancient Syria and Mesopotamia, where it also served as an economic and political status symbol.
Cuneiform Cuneiform is a logo-syllabic writing system, script that was used to write several languages of the Ancient Near East, Ancient Middle East. The script was in active use from the early Bronze Age until the beginning of the Common Era. It is nam ...
writings from as early as the mid-third millennium
BCE Common Era (CE) and Before the Common Era (BCE) are year notations for the Gregorian calendar (and its predecessor, the Julian calendar), the world's most widely used calendar era. Common Era and Before the Common Era are alternatives to the o ...
describe the animal as a hybrid but do not provide the precise taxonomical nature of the breeding that produced it. Modern
paleogenomics Paleogenomics is a field of science based on the reconstruction and analysis of genomic information in extinct species. Improved methods for the extraction of ancient DNA (aDNA) from museum artifacts, ice cores, archeological or paleontological site ...
has revealed it to have been the offspring of a female domesticated
donkey The domestic donkey is a hoofed mammal in the family Equidae, the same family as the horse. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a ...
and a wild male
Syrian wild ass The Syrian wild ass (''Equus hemionus hemippus''), less commonly known as a hemippe, an achdari, or a Mesopotamian or Syrian onager, is an extinct subspecies of onager native to the Arabian peninsula and surrounding areas. It ranged across present ...
. They fell out of favor after the introduction of domesticated horses into the region at the end of the 3rd millennium BCE.


Elite equids

Third-millennium BCE cuneiform from the kingdom of
Ebla Ebla ( Sumerian: ''eb₂-la'', ar, إبلا, modern: , Tell Mardikh) was one of the earliest kingdoms in Syria. Its remains constitute a tell located about southwest of Aleppo near the village of Mardikh. Ebla was an important center t ...
and the Mesopotamian region of Diyala name several types of equids (''ANŠE'', ), including one specified as the ''kúnga'' (''ANŠE BAR.AN'', ), which appear between about 2600 and 2000 BCE. These expensive animals, highly valued by the elite, were purpose-bred at Nagar, the rulers of which used them themselves and monopolized their production for distribution in the region. Records from Ebla report repeated expensive purchases of kunga equids from Nagar, and it was apparently in relation to this trade that the 'high superintendents of charioteers' and those responsible for maintaining the Ebla kunga herd traveled to Nagar. The Ebla king gave them as gifts to other rulers. It has been suggested that the kunga trade was central to the economies of the region's kingdoms, and that the ostentatious display of such expensive animals in official art directly associated them with kingship and power. A pair of seals from the period, including one from Nagar, depict equids with gods in the divine realm.


Hybrid nature

Contemporary descriptions of the production of the kunga seem to indicate that they were hybrids, and there are indications that, like most hybrid equids, they were sterile. For example, foals are described in nursery herds with adult donkeys or onagers and donkey foals, never with kunga parents. Production would thus have been an intensive process: they would not have established a domesticated line, but rather each individual kunga had to be produced ''de novo'' by breeding two parental species anew, without the opportunity for improvement through selective breeding. Likewise, the necessity of repeated purchase of new animals from their limited production centers to maintain a stable of kunga suggests they could not be bred.


Depictions

Kunga were used as draft animals, with smaller males and females used for pulling plows, while 'superior' males are described in more ceremonial and martial roles, pulling the four-wheeled battle wagons and chariots of kings and gods. Equids appear in this role in official imagery such as the ca. 2600 BCE
Standard of Ur The Standard of Ur is a Sumerian artifact of the 3rd millennium BC that is now in the collection of the British Museum. It comprises a hollow wooden box measuring wide by long, inlaid with a mosaic of shell, red limestone and lapis lazuli. It ...
mozaic and numerous surviving seals, while a rein ring similar to those depected in the mozaic has been found at Ur, decorated with an equid. These depictions are likely kunga rather than donkeys, which appear only in lesser roles in descriptions. Illustrations appear to show the draft team of equids being controlled by strings passed through rings placed in the equids' upper lips. Their appearance in formal administrative cuneiform and official art seems to parallel the contemporary development of kingships in the region, suggesting a propagandistic association of the kunga with royalty.


Archaeology and paleogenomics

They are known to have been used for funerary purposes, as demonstrated by high-status-funeral disbursement records for harnesses, and they have been identified with more than 40 equids that were sacrificed and ceremonially buried in elite graves at
Umm el-Marra Umm el-Marra, ( ar, أم المرى), east of modern Aleppo in the Jabbul Plain of northern Syria, was one of the ancient Near East's oldest cities, located on a crossroads of two trade routes northwest of Ebla, in a landscape that was much more f ...
, Syria, in separate chambers from the burials of adult humans but many accompanied by human infants with signs of having been sacrificed. These buried kunga may have been intended either as offerings to deities, or as companions of the buried human elites, while such burials may also have served a legitimizing role for the royal lines and elite, with sacrificed 'royal' equids serving as analogs of human royals. Like the 'superior' kunga of cuneiform, these equids were all male, ranging in size from 1.19 m to 1.36 m. There are inherent challenges in identifying the species of equid skeletons, but the Umm el-Marra equids shared signs of domestication such as bit wear and evidence of
fodder Fodder (), also called provender (), is any agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food given to the animals (includin ...
ing rather than
grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and other ...
. They had a prominent overbite that suggested all represented a common population of equid, while their bones had a combination of
onager The onager (; ''Equus hemionus'' ), A new species called the kiang (''E. kiang''), a Tibetan relative, was previously considered to be a subspecies of the onager as ''E. hemionus kiang'', but recent molecular studies indicate it to be a distinct ...
and donkey characteristics, being sized more like the former, but with the greater robustness of the latter, as might be expected in a hybrid between the two equid species. Such a hybrid would have been stronger and faster than the donkey, while less intractable to taming than the Syrian wild ass. Their hypothesized taxonomic identity was proven by a genomic analysis reported in 2022 that compared genomes from several of these skeletons with those of extant and extinct equids, and concluded that all of the Umm el-Marra skeletons were
F1 hybrid An F1 hybrid (also known as filial 1 hybrid) is the first filial generation of offspring of distinctly different parental types. F1 hybrids are used in genetics, and in selective breeding, where the term F1 crossbreed may be used. The term is somet ...
progeny of captured male Syrian wild asses with female domesticated donkeys (jenny). These results make the kunga the earliest known human-engineered hybrid animal, predating the earliest
mule The mule is a domestic equine hybrid between a donkey and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two po ...
by about 1500 years. The preference for a jenny over a jack (male) as the donkey parent represents a conscious choice to have the more tractable domestic species as the maternal parent for simpler husbandry. That all tested individuals were F1 hybrids reinforces the likelihood that kunga were sterile.


Fate

Though the kunga held its elite status for half a millennium, it would be supplanted by domestic horses, introduced to the region at the end of the third millennium BCE and after that time seen filling the roles previously occupied by the kunga, which rapidly disappear from the historical record. A similar hybrid was reportedly produced at the
London Zoo London Zoo, also known as ZSL London Zoo or London Zoological Gardens is the world's oldest scientific zoo. It was opened in London on 27 April 1828, and was originally intended to be used as a collection for science, scientific study. In 1831 o ...
in 1883, but the subsequent extinction of the Syrian wild ass makes it impossible now to reproduce the kunga's precise taxonomic cross.


References

{{taxonbar, from =Q110811351 Ancient Mesopotamia Equid hybrids Livestock Extinct mammals