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''Kalmia latifolia'', the mountain laurel, calico-bush, or spoonwood, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
in the heath family
Ericaceae The Ericaceae are a family of flowering plants, commonly known as the heath or heather family, found most commonly in acidic and infertile growing conditions. The family is large, with c.4250 known species spread across 124 genera, making it th ...
, that is native to the eastern United States. Its range stretches from southern
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and nor ...
south to northern
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
, and west to
Indiana Indiana () is a U.S. state in the Midwestern United States. It is the 38th-largest by area and the 17th-most populous of the 50 States. Its capital and largest city is Indianapolis. Indiana was admitted to the United States as the 19th ...
and
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is bord ...
. Mountain laurel is the state flower of
Connecticut Connecticut () is the southernmost state in the New England region of the Northeastern United States. It is bordered by Rhode Island to the east, Massachusetts to the north, New York (state), New York to the west, and Long Island Sound to the ...
and
Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...
. It is the namesake of Laurel County in Kentucky, the city of Laurel, Mississippi, and the
Laurel Highlands The Laurel Highlands is a region in southwestern Pennsylvania made up of Fayette County, Somerset County and Westmoreland County. It has a population of about 600,000 people. The region is approximately fifty-five miles southeast of Pit ...
in southwestern Pennsylvania.


Growth

''Kalmia latifolia'' is an
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
shrub growing tall. The leaves are 3–12 cm long and 1–4 cm wide. The flowers are hexagonal, sometimes appearing to be pentagonal, ranging from light pink to white, and occur in clusters. There are several named
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s that have darker shades of pink, red and maroon. It blooms in May and June. All parts of the plant are poisonous. The roots are fibrous and matted. The plant is naturally found on rocky slopes and mountainous forest areas. It thrives in acid soil, preferring a
soil pH Soil pH is a measure of the acidity or basicity (alkalinity) of a soil. Soil pH is a key characteristic that can be used to make informative analysis both qualitative and quantitatively regarding soil characteristics. pH is defined as the ne ...
in the 4.5 to 5.5 range. The plant often grows in large thickets, covering great areas of forest floor. In the
Appalachians The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. They ...
, it can become a tree but is a shrub farther north. The species is a frequent component of oak-heath forests. In low, wet areas it grows densely, but in dry uplands has a more sparse form. In the southern Appalachians, laurel thickets are referred to as "laurel hells" because it is nearly impossible to pass through one.


Reproduction

''Kalmia latifolia'' is notable for its unusual method of dispensing its pollen. As the flower grows, the filaments of its
stamens The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
are bent and brought into tension. When an insect lands on the flower, the tension is released, catapulting the pollen forcefully onto the insect. Experiments have shown the flower capable of flinging its pollen up to 15 cm. Physicist Lyman J. Briggs became fascinated with this phenomenon in the 1950s after his retirement from the
National Bureau of Standards The National Institute of Standards and Technology (NIST) is an agency of the United States Department of Commerce whose mission is to promote American innovation and industrial competitiveness. NIST's activities are organized into physical sci ...
and conducted a series of experiments in order to explain it.


Etymology

''Kalmia latifolia'' is also known as ivybush or spoonwood (because Native Americans used to make their spoons out of it). The plant was first recorded in America in 1624, but it was named after the Finnish explorer and botanist
Pehr Kalm Pehr Kalm (6 March 1716 – 16 November 1779), also known as Peter Kalm, was a Swedish explorer, botanist, naturalist, and agricultural economist. He was one of the most important apostles of Carl Linnaeus. In 1747, he was commissioned by th ...
(1716–1779), who sent samples to
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
. The Latin
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
''latifolia'' means "with broad leaves" – as opposed to its sister species ''
Kalmia angustifolia ''Kalmia angustifolia'' is a flowering shrub in the family Ericaceae, commonly known as sheep laurel. It is distributed in eastern North America from Ontario and Quebec south to Virginia. It grows commonly in dry habitats in the boreal forest, a ...
'', "with narrow leaves". Despite the name "mountain laurel", ''Kalmia latifolia'' is not closely related to the true laurels of the family
Lauraceae Lauraceae, or the laurels, is a plant family that includes the true laurel and its closest relatives. This family comprises about 2850 known species in about 45 genera worldwide (Christenhusz & Byng 2016 ). They are dicotyledons, and occur m ...
.


Cultivation

The plant was originally brought to Europe as an
ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that ...
during the 18th century. It is still widely grown for its attractive flowers and year-round evergreen leaves. Elliptic, alternate, leathery, glossy evergreen leaves (to 5" long) are dark green above and yellow green beneath and reminiscent of the leaves of rhododendrons. All parts of this plant are toxic if ingested. Numerous
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s have been selected with varying flower color. Many of the cultivars have originated from the Connecticut Experiment Station in Hamden and from the plant breeding of Dr. Richard Jaynes. Jaynes has numerous named varieties that he has created and is considered the world's authority on ''Kalmia latifolia''. In the UK the following cultivars have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
: *'Freckles' – pale pink flowers, heavily spotted *'Little Linda' – dwarf cultivar to *'Olympic Fire' – red buds opening pale pink *'Pink Charm'


Ecology

''Kalmia latifolia'' has been marked as a pollinator plant, supporting and attracting
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises ...
and
hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With about 361 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are ...
s.


Wood

The wood of the mountain laurel is heavy and strong but brittle, with a close, straight grain. It has never been a viable commercial crop as it does not grow large enough, yet it is suitable for wreaths, furniture, bowls and other household items. It was used in the early 19th century in wooden-works clocks. Root
burl A burl (American English) or burr (British English) is a tree growth in which the grain has grown in a deformed manner. It is commonly found in the form of a rounded outgrowth on a tree trunk or branch that is filled with small knots from dor ...
s were used for pipe bowls in place of imported briar burls unattainable during
World War II World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the World War II by country, vast majority of the world's countries—including all of the great power ...
. It can be used for handrails or guard rails.


Toxicity

Mountain laurel is
poison Poison is a chemical substance that has a detrimental effect to life. The term is used in a wide range of scientific fields and industries, where it is often specifically defined. It may also be applied colloquially or figuratively, with a broa ...
ous to several animals, including
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
s,
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
s,
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Adult females are referred to as cows and adult ...
,
deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the re ...
, monkeys, and humans, due to grayanotoxin and
arbutin Arbutin is a glycoside; a glycosylated hydroquinone extracted from the bearberry plant in the genus ''Arctostaphylos'' among many other medicinal plants, primarily in the family Ericaceae. Applied topically, it inhibits tyrosinase and thus prev ...
. The green parts of the plant,
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s, twigs, and
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametop ...
are all toxic, including food products made from them, such as toxic honey that may produce neurotoxic and
gastrointestinal The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The GI tract contains all the major organs of the digestive system, in humans and ...
symptoms in humans eating more than a modest amount. Symptoms of
toxicity Toxicity is the degree to which a chemical substance or a particular mixture of substances can damage an organism. Toxicity can refer to the effect on a whole organism, such as an animal, bacterium, or plant, as well as the effect on a subs ...
begin to appear about 6 hours following ingestion. Symptoms include irregular or difficulty breathing, anorexia, repeated swallowing, profuse
salivation Saliva (commonly referred to as spit) is an extracellular fluid produced and secreted by salivary glands in the mouth. In humans, saliva is around 99% water, plus electrolytes, mucus, white blood cells, epithelial cells (from which DNA can ...
, watering of the eyes and nose, cardiac distress, incoordination, depression,
vomiting Vomiting (also known as emesis and throwing up) is the involuntary, forceful expulsion of the contents of one's stomach through the mouth and sometimes the nose. Vomiting can be the result of ailments like food poisoning, gastroenter ...
, frequent
defecation Defecation (or defaecation) follows digestion, and is a necessary process by which organisms eliminate a solid, semisolid, or liquid waste material known as feces from the digestive tract via the anus. The act has a variety of names ranging f ...
, weakness, convulsions, paralysis,
coma A coma is a deep state of prolonged unconsciousness in which a person cannot be awakened, fails to respond normally to painful stimuli, light, or sound, lacks a normal wake-sleep cycle and does not initiate voluntary actions. Coma patients exhi ...
, and eventually death.
Necropsy An autopsy (post-mortem examination, obduction, necropsy, or autopsia cadaverum) is a surgical procedure that consists of a thorough examination of a corpse by dissection to determine the cause, mode, and manner of death or to evaluate any di ...
of animals who have died from spoonwood poisoning show gastrointestinal
hemorrhage Bleeding, hemorrhage, haemorrhage or blood loss, is blood escaping from the circulatory system from damaged blood vessels. Bleeding can occur internally, or externally either through a natural opening such as the mouth, nose, ear, urethra, v ...
.


Use by Native Americans

The
Cherokee The Cherokee (; chr, ᎠᏂᏴᏫᏯᎢ, translit=Aniyvwiyaʔi or Anigiduwagi, or chr, ᏣᎳᎩ, links=no, translit=Tsalagi) are one of the indigenous peoples of the Southeastern Woodlands of the United States. Prior to the 18th century, th ...
use the plant as an
analgesic An analgesic drug, also called simply an analgesic (American English), analgaesic (British English), pain reliever, or painkiller, is any member of the group of drugs used to achieve relief from pain (that is, analgesia or pain management). It ...
, placing an
infusion Infusion is the process of extracting chemical compounds or flavors from plant material in a solvent such as water, oil or alcohol, by allowing the material to remain suspended in the solvent over time (a process often called steeping). An in ...
of leaves on scratches made over location of the pain. They also rub the bristly edges of ten to twelve leaves over the skin for
rheumatism Rheumatism or rheumatic disorders are conditions causing chronic, often intermittent pain affecting the joints or connective tissue. Rheumatism does not designate any specific disorder, but covers at least 200 different conditions, including ar ...
, crush the leaves to rub brier scratches, use an infusion as a wash "to get rid of pests", use a compound as a
liniment Liniment (from la, linere, meaning "to anoint"), also called embrocation and heat rub, is a medicated topical preparation for application to the skin. Some liniments have viscosity similar to that of water; others are lotion or balm; still oth ...
, rub leaf ooze into the scratched skin of ball players to prevent cramps, and use a leaf salve for healing. They also use the wood for carving.Hamel, Paul B. and Mary U. Chiltoskey 1975 Cherokee Plants and Their Uses – A 400 Year History. Sylva, N.C. Herald Publishing Co. (p. 42)


Gallery

File:Mountain Laurel Kalmia latifolia 'Olympic Wedding' Leaves and Buds 2575px.jpg, ''K. latifolia'' leaves and early buds File:Buberel unknown flower 12.jpg, Flower buds File:Kalmia latifolia species.jpg, Beginning to bloom File:Mountain Loral Flowers.jpg, Full bloom File:Mountain Laurel Kalmia latifolia 'Olympic Wedding' Young Old Flowers 3264px.jpg, Blooming and wilted flowers on the same flower head File:Bee on a mountain laurel on Occoneechee Mountain.jpg, Bee pollinating mountain laurel on Occoneechee Mountain


References


External links


Connecticut Botanical Society Profile: ''Kalmia latifolia''
{{Taxonbar, from=Q1235131 latifolia Flora of the Appalachian Mountains Trees of the Northeastern United States Trees of the Southeastern United States Trees of the Great Lakes region (North America) Natural history of the Great Smoky Mountains Plants used in traditional Native American medicine Plants described in 1753 Taxa named by Carl Linnaeus Symbols of Connecticut Symbols of Pennsylvania Garden plants of North America